2009 in paleontology
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Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils.[1] This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2009.
Arthropods
Cephalopods
Three new species of extinct Octopoda discovered in 2009. The species – Keuppia hyperbolaris, Keuppia levante, and Styletoctopus annae – lived about 95 million years ago, and bear a strong resemblance to modern octopuses, suggesting that the Octopoda order has remained relatively unchanged for tens of millions of years. The fossils included evidence of arms, muscles, rows of suckers, ink, and internal gills. The discovery was made by a team led by Dirk Fuchs of the Freie University, which is located at Berlin, Germany.[2] The fossils were found at Hakel and Hadjoula, Lebanon.[3] Various new ammonoid taxa were named, including Ivoites.
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Gen et sp nov |
Valid |
Fuchs, Bracchi, & Weis |
Upper Cenomanian |
|||||
gen et sp nov |
Valid |
Fuchs, Bracchi, & Weis |
Upper Cenomanian |
|||||
gen nov |
Valid |
De Baets, Klug, & Korn |
Lower Emsian |
Cartilaginous fish
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Gen. et sp. nov |
Valid |
Wang et al. |
Late Permian |
A member of Hybodontoidea. Genus includes new species G. tungshengi. |
||||
Gen. et sp. nov |
Disputed |
Kaddumi |
Late Cretaceous (Maastrichtian) |
A member of the family Sclerorhynchidae. The type species is H. minutadens. |
||||
Gen. et sp. nov |
Valid |
|
A member of Iniopterygidae. The type species is P. stahlae. |
|||||
Gen. et sp. nov |
Valid |
|
A member of Iniopterygidae. The type species is R. zangerli. |
Bony fish
Newly named bony fish | |||||||
---|---|---|---|---|---|---|---|
Name | Status | Authors | Age | Unit | Location | Notes | Images |
Aphanius yerevanicus[9] |
Valid |
|
Late Miocene |
A pupfish, a species of Aphanius. | |||
Carpathoserranoides[10] |
Valid |
|
A member of Percoidei. The type species is C. brnoensis; genus also includes C. polonicus. | ||||
Caucasoserranoides[10] |
Valid |
|
A member of Percoidei. The type species is C. morozkiensis. | ||||
Enchodus harranaensis[11] |
Disputed |
|
Late Cretaceous (late Maastrichtian) |
||||
Eophryne[12] |
Valid |
|
Eocene (late Ypresian) |
Monte Bolca locality |
A frogfish. The type species is Eophryne barbutii. | ||
Gogosardina[13] |
Valid |
|
Late Devonian |
An early ray-finned fish. Genus includes new species G. coatesi. | |||
Heddleichthys[14] |
Valid |
|
Famennian (Late Devonian) |
Dura Den Formation |
|||
Hendrixella[15] |
Valid |
|
Eocene (late Ypresian) |
Monte Bolca locality |
A member of Percoidei of uncertain phylogenetic placement. The type species is Hendrixella grandei. | ||
Langlieria[16] |
Valid |
|
Famennian (Late Devonian) |
Evieux Formation |
|||
Oligoserranoides[10] |
Valid |
|
Abkhazia |
A member of Percoidei. The type species is "Smerdis" budensis Heckel (1856); genus also includes "Serranus" comparabilis Daniltshenko (1960). | |||
Postredectes[17] |
Disputed |
|
Late Cretaceous (late Maastrichtian) |
Muwaqqar Chalk Marl Formation |
A member of the family Ichthyodectidae. The type species is P. harranaensis. | ||
Ridewoodichthys[18] |
Valid |
|
Early Paleocene |
An arowana; a new genus for "Brychaetus" caheni Taverne (1969). | |||
Saurocephalus longicorpus[19] |
Disputed |
|
Late Cretaceous (late Maastrichtian) |
Muwaqqar Chalk Marl Formation |
|||
Ungarnia[10] |
Valid |
|
A member of Percoidei. The type species is "Serranus" transsylvanicus Bohm (1942). |
- Shimada, K.; Everhart, M.J. (2009). "First record of Anomoeodus (Osteichthyes: Pycnodontiformes) from the Upper Cretaceous Niobrara Chalk of western Kansas". Transactions of the Kansas Academy of Science. 112 (1/2): 98–102. doi:10.1660/062.112.0212.
Amphibians
Newly named amphibians
Name | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|
Valid |
|
Late Carboniferous |
|||||
Valid |
|
Lower Cretaceous |
Possible stem neobatrachian |
||||
Valid |
|
Lower Cretaceous |
A hyloid | ||||
Valid |
|
The smallest known ostodolepid microsaur | |||||
Valid |
|
Turonian |
|||||
Valid |
|
Early Cretaceous |
A cryptobranchoid salamander | ||||
Valid |
|
Early Permian |
Basal reptiles
Newly named basal reptiles
Name | Status | Authors | Discovery year | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Valid |
|
A basal parareptile | ||||||
Valid |
|
early Late Olenekian |
Turtles
Newly named turtles
Name | Status | Authors | Discovery year | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Valid |
|
Turonian (Late Cretaceous) |
||||||
Valid |
|
Late Cretaceous |
||||||
Valid |
|
Late Jurassic/Lower Cretaceous |
||||||
Valid |
|
Paleocene |
USA |
New genus for "Plesiobaena" putorius Gaffney, 1972 | ||||
Valid |
|
Holocene |
A tortoise. | |||||
Valid |
|
A basal member of Testudinata. The type species is C. tenertesta. The genus Chinlechelys was considered to be a junior synonym of the genus Proganochelys by Joyce (2017), though the author maintained C. tenertesta as a distinct species within the latter genus.[34] | ||||||
Junior synonym |
|
Early Paleocene |
USA |
Junior synonym of Hutchemys.[36] | ||||
Valid |
|
Middle Jurassic |
||||||
Valid |
|
Paleocene |
USA |
A plastomenine softshell turtles | ||||
Valid |
Late Cretaceous |
|||||||
Valid |
|
Maastrichtian |
USA |
A baenid | ||||
Valid |
|
Late Cretaceous |
USA |
A baenid | ||||
Junior synonym |
|
Early Paleocene |
USA |
Junior synonym of Hutchemys.[36] |
Archosauromorphs
Basal archosauromorphs
Newly named basal archosauromorphs | |||||||
---|---|---|---|---|---|---|---|
Name | Status | Authors | Age | Unit | Location | Notes | Images |
Valid |
|
earliest Late Olenekian |
A long−necked archosauromorph |
Archosaurs
Lepidosauromorphs
Basal lepidosauromorphs
Newly named basal lepidosauromorphs | |||||||
---|---|---|---|---|---|---|---|
Name | Status | Authors | Age | Unit | Location | Notes | Images |
Valid |
Early Olenekian |
A basal kuehneosaurid |
|||||
Valid |
|
earliest Late Olenekian |
A basal lepidosauromorph |
Plesiosaurs
- In 2009, in Svalbard, Norway a new pliosaur was found by Jorn Hurum. It currently is codenamed as "Predator X."
- O'Keefe, F. R.; Street, H. P. (2009). "Osteology of the cryptocleidoid plesiosaur Tatenectes laramiensis, with comments on the taxonomic status of the Cimoliasauridae". Journal of Vertebrate Paleontology. 29 (1): 48–57. doi:10.1671/039.029.0118.
Newly named plesiosaurs | |||||||
---|---|---|---|---|---|---|---|
Name | Status | Authors | Age | Unit | Location | Notes | Images |
Valid |
|
Late Jurassic (Oxfordian) |
|||||
Valid |
|
Early Cretaceous (Albian) |
Replacement name for Nichollsia Druckenmiller & Russell, 2008, preoccupied by an isopod genus Nichollsia Chopra & Tiwari, 1950 |
||||
Disputed |
|
Late Cretaceous (Maastrichtian) |
A member of Polycotylidae. The type species is R. singularis. |
Squamates
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
Valid |
Schulp, Bardet & Bouya |
A mosasaur. |
|||||
Sp. nov |
Disputed |
Kaddumi |
Late Cretaceous (Maastrichtian) |
A mosasaur. |
||||
Gen. et sp. nov |
Disputed |
Kaddumi |
Late Cretaceous (Maastrichtian) |
Muwaqqar Chalk Marl Formation |
A globidensine mosasaur. The type species is H. khuludae. | |||
Sp. nov |
Disputed |
Kaddumi |
Late Cretaceous (Maastrichtian) |
A mosasaur. |
||||
Sp. nov |
Disputed |
Kaddumi |
Late Cretaceous (Maastrichtian) |
A mosasaur. | ||||
Tenerasaurus[52] |
Gen. et sp. nov |
Junior synonym |
Kaddumi |
Late Cretaceous (Maastrichtian) |
A mosasaur. The type species is T. hashimi. /> Lindgren, Kaddumi & Polcyn (2013) reinterpreted the holotype specimen as belonging to a member of the genus Prognathodon of uncertain specific assignment.[53] |
|||
Gen. et sp. nov |
Valid |
Head et al. |
Paleocene |
Cerrejón Formation |
In February, the fossils of 28 individual T. cerrejonensis (Titanoboas) were announced to have been found in the coal mines of Cerrejón, La Guajira, Colombia.[55] |
Synapsids
Non-mammalian
Name | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|
Valid |
|
Middle Triassic (Ladinian) |
|||||
Valid |
|
Middle Permian (Roadian) |
|||||
Valid |
|
Late Jurassic (Oxfordian) |
Mammals
Plants
Angiosperms
Name | Novelty | Status | Authors | Age | Unit | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
gen et sp nov |
Valid |
Coiffard et al. |
uppermost Albian-lowermost Cenomanian |
Charente-Maritimes |
Relevant research in other sciences
Evolutionary biology
- A study is published that proposes that females from certain taxa use ornaments as a criterion for mate choice because other dimorphic structures, like biological "weaponry" could be used to coerce or force them to mate.[60]
- A study concludes that biotic factors have more pronounced local and short term evolutionary impacts than abiotic factors, which in turn have a more pronounced effect through time and on biodiversity as a whole.[61]
Extinction
A study noting the effects of the KT mass extinction on Earth's modern biota is published.[62]
Geology
- Zhang, H.; Wei, Z.-L.; Liu, X.-M.; Li, D. (2009). "Constraints on the age of the Tuchengzi Formation by LA-ICP-MS dating in northern Hebei-western Liaoning, China". Science China Earth Sciences. 52 (4): 461–470. doi:10.1007/s11430-009-0052-9.
Ichnology
- Bedatou E., Melchor R.N., Genise J.F. (2009). "Complex palaeosol ichnofabrics from Late Jurassic-Early Cretaceous volcaniclastic successions of central Patagonia, Argentina". Sedimentary Geology. 218 (1–4): 74–102. doi:10.1016/j.sedgeo.2009.04.005.
{{cite journal}}
: CS1 maint: multiple names: authors list (link)
Paleobiogeography
- Pereda-Suberbiola X (2009). "Biogeographical affinities of Late Cretaceous continental tetrapods of Europe: a review". Bulletin de la Société Géologique de France. 180 (1): 57–71. doi:10.2113/gssgfbull.180.1.57.
Paleoecology
- Nicolas M., Rubidge B.S. (2009). "Changes in Permo-Triassic terrestrial tetrapod ecological representation in the Beaufort Group (Karoo Supergroup) of South Africa". Lethaia. 43: 45–59. doi:10.1111/j.1502-3931.2009.00171.x.
Footnotes
Complete author list
As science becomes more collaborative, papers with large numbers of authors are becoming more common. To prevent the deformation of the tables, these footnotes list the contributors to papers that erect new genera and have many authors.
References
- ^ Gini-Newman, Garfield; Graham, Elizabeth (2001). Echoes from the past: world history to the 16th century. Toronto: McGraw-Hill Ryerson Ltd. ISBN 9780070887398. OCLC 46769716.
- ^ Rare fossil octopuses found, NBC News, March 18, 2009
- ^ New Octopus from the late Cretaceous of Hakel and Hadjoula, Lebanon, Palaeontology, Volume 52, Issue 1, Pages 65-81
- ^ a b Fuchs, D.; Bracchi, G.; Weis, R. (2009). "New Octopods (Cephalopoda: Coleoidea) from the Late Cretaceous (Upper Cenomanian) of Hakel and Hadjoula, Lebanon". Palaeontology. 52 (1): 65–81. doi:10.1111/j.1475-4983.2008.00828.x. Archived from the original on 14 March 2011. Retrieved 24 March 2010.
- ^ De Baets, K.; Klug, C.; Korn, D. (2009). "Anetoceratinae (Ammonoidea, Early Devonian) from the Eifel and Harz Mountains (Germany), with a revision of their genera". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 252 (3): 361–376. doi:10.1127/0077-7749/2009/0252-0361.
- ^ N.‐Z. Wang; X. Zhang; M. Zhu; W.‐J. Zhao (2009). "A new articulated hybodontoid from Late Permian of northwestern China". Acta Zoologica. 90 (s1): 159–170. doi:10.1111/j.1463-6395.2008.00382.x.
- ^ Hani Faig Kaddumi (2009). "A new genus and species of sawfishes (Chondrichthyes: Batoidea) from the late Maastrichtian sediments of Harrana". In Hani Faig Kaddumi (ed.). Fossils of the Harrana Fauna and the Adjacent Areas. pp. 178–187.
- ^ a b Eileen D. Grogan; Richard Lund (2009). "Two new iniopterygians (Chondrichthyes) from the Mississippian (Serpukhovian) Bear Gulch Limestone of Montana with evidence of a new form of chondrichthyan neurocranium". Acta Zoologica. 90 (s1): 134–151. doi:10.1111/j.1463-6395.2008.00371.x.
- ^ Davit Vasilyan; Bettina Reichenbacher; Giorgio Carnevale (2009). "A new fossil Aphanius species from the Upper Miocene of Armenia (Eastern Paratethys)". Paläontologische Zeitschrift. 83 (4): 511–519. doi:10.1007/s12542-009-0034-4.
- ^ a b c d A.M. Prokofiev (2009). "К систематике олигоценовых перкоидных рыб, известных под сборным названием "Serranus budensis", с выделением новых таксонов". Актуальные проблемы современной науки. 46: 199–222.
- ^ Hani Faig Kaddumi (2009). "A new species of large Enchodus fishes (Aulopiformes: Enchodontidae) from the late Maastrichtian of Harrana". In Hani Faig Kaddumi (ed.). Fossils of the Harrana Fauna and the Adjacent Areas. pp. 204–214.
- ^ Giorgio Carnevale; Theodore W. Pietsch (2009). "An Eocene frogfish from Monte Bolca, Italy: the earliest known skeletal record for the family". Palaeontology. 52 (4): 745–752. doi:10.1111/j.1475-4983.2009.00874.x.
- ^ Brian Choo; John A. Long; Katherine Trinajstic (2009). "A new genus and species of basal actinopterygian fish from the Upper Devonian Gogo Formation of Western Australia". Acta Zoologica. 90 (s1): 194–210. doi:10.1111/j.1463-6395.2008.00370.x.
- ^ Snitting, D. (2009). "Heddleichthys- a new tristichopterid genus from the Dura Den Formation, Midland Valley, Scotland (Famennian, Late Devonian)". Acta Zoologica. 90: 273–284. doi:10.1111/j.1463-6395.2008.00376.x.
- ^ Alexandre F. Bannikov; Giorgio Carnevale (2009). "A new percoid fish from the Eocene of Monte Bolca, Italy: Hendrixella grandei gen. & sp. nov". Swiss Journal of Geosciences. 102 (3): 481–488. doi:10.1007/s00015-009-1331-3.
- ^ Clément, G; Snitting, D; Ahlberg, PE; Gaël Clément; Daniel Snitting; Per Erik Ahlberg (July 2009). "A new Tristichopterid (Sarcopterygii, Tetrapodomorpha) from the Evieux Formation (Upper Devonian) of Belgium". Palaeontology. 52 (4): 823–836. doi:10.1111/j.1475-4983.2009.00876.x.
- ^ Hani Faig Kaddumi (2009). "Ichthyodectids of the late Maastrichtian sediments of the Muwaqqar Chalk Marl Formation of Harrana". In Hani Faig Kaddumi (ed.). Fossils of the Harrana Fauna and the Adjacent Areas. pp. 232–239.
- ^ Louis Taverne (2009). "Ridewoodichthys, a new genus for Brychaetus caheni from the marine Paleocene of Cabinda (Africa): re-description and comments on its relationships within the Osteoglossidae (Teleostei, Osteoglossomorpha)". Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Sciences de la Terre. 79: 147–153.
- ^ Hani Faig Kaddumi (2009). "Saurodontids (Ichthyodectiformes: Saurocephalus) of Harrana with a description of a new species from the late Maastrichtian Muwaqqar Chalk Marl Formation". In Hani Faig Kaddumi (ed.). Fossils of the Harrana Fauna and the Adjacent Areas. pp. 215–231.
- ^ Angela C. Milner; Andrew R. Milner; Stig A. Walsh (2009). "A new specimen of Baphetes from Nýřany, Czech Republic and the intrinsic relationships of the Baphetidae". Acta Zoologica. 90 (s1): 318–334. doi:10.1111/j.1463-6395.2008.00340.x.
- ^ a b Báez, Ana M.; Moura, Geraldo J.B.; Gómez, Raúl O.; Ana M. Báez; Geraldo J.B. Moura; Raúl O. Gómez (2009). "Anurans from the Lower Cretaceous Crato Formation of northeastern Brazil: implications for the early divergence of neobatrachians". Cretaceous Research. 30 (4): 829–846. doi:10.1016/j.cretres.2009.01.002.
- ^ Anderson, J.S.; Scott, D.; Reisz, R.R. (2009). "Nannaroter mckinziei, a new ostodolepid 'microsaur' (Tetrapoda, Lepospondyli, Recumbirostra) from the Early Permian of Richards Spur (Ft. Sill), Oklahoma". Journal of Vertebrate Paleontology. 29 (2): 379–388. doi:10.1671/039.029.0222.
- ^ Skutschas, Pavel P.; Pavel P. Skutschas (2009). "Re-Evaluation of Mynbulakia Nesov, 1981 (Lissamphibia: Caudata) and Description of a New Salamander Genus from the Late Cretaceous of Uzbekistan". Journal of Vertebrate Paleontology. 29 (3): 659–664. doi:10.1671/039.029.0326.
- ^ Zhang, Guilin; Wang, Yuan; Jones, Marc E.H.; Evans, Susan E.; Guilin Zhang; Yuan Wang; Marc E.H. Jones; Susan E. Evans (2009). "A new Early Cretaceous salamander (Regalerpeton weichangensis gen. et sp. nov.) from the Huajiying Formation of northeastern China". Cretaceous Research. 30 (3): 551–558. doi:10.1016/j.cretres.2008.10.004.
- ^ Klembara, Jozef (2009). "The skeletal anatomy and relationships of a new discosauriscid seymouriamorph from the lower Permian of Moravia (Czech Republic)". Annals of Carnegie Museum. 77 (4): 451–483. doi:10.2992/0097-4463-77.4.451.
- ^ Sean P. Modesto; Diane M. Scott; Robert R. Reisz (2009). "A new parareptile with temporal fenestration from the Middle Permian of South Africa". Canadian Journal of Earth Sciences. 46 (1): 9–20. Bibcode:2009CaJES..46....9M. doi:10.1139/E09-001. Retrieved 14 August 2010.[permanent dead link]
- ^ Magdalena Borsuk−Białynicka; Mariusz Lubka (2009). "Procolophonids from the Early Triassic of Poland" (PDF). Paleontologica Polonica. 65: 107–144. Retrieved 12 August 2010..
- ^ Octávio Mateus; Louis Jacobs; Michael Polcyn; Anne S. Schulp; Diana Vineyard; André Buta Neto; Miguel Telles Antunes (2009). "The oldest African eucryptodiran turtle from the Cretaceous of Angola" (PDF). Acta Palaeontologica Polonica. 54 (4): 581–588. doi:10.4202/app.2008.0063. Retrieved 21 June 2010.
- ^ Vandermark, D.; Tarduno, J.A.; Brinkman, D.B.; Cottrell, R.D.; Mason, S. (2009). "New Late Cretaceous macrobaenid turtle with Asian affinities from the High Canadian Arctic: dispersal via ice-free polar routes". Geology. 37 (2): 183–186. Bibcode:2009Geo....37..183V. doi:10.1130/G25415A.1.
- ^ Tong, H.; Claude, J.; Naksri, W.; Suteethorn, V.; Buffetaut, E.; Khansubha, S.; Wongko, K. & Yuangdetkla, P. (2009). "Basilochelys macrobios n. gen. and n. sp., a large cryptodiran turtle from the Phu Kradung Formation (latest Jurassic-earliest Cretaceous) of the Khorat Plateau, NE Thailand". In: Buffetaut, E.; Cuny, G.; Le Loeuff, J. & Suteethorn, V. (eds.). Late Palaeozoic and Mesozoic Ecosystems in SE Asia. Geological Society, London, Special Publications 315: 229-243.
- ^ a b Lyson, T.R.; Joyce, W.G. (2009). "A revision of Plesiobaena (Testudinoes: Baenidae) and an assessment of Baenid ecology across the K/T boundary". Journal of Paleontology. 83 (6): 833–853. doi:10.1666/09-035.1.
- ^ Richard Franz; Shelley E. Franz (2009). "A new fossil land tortoise in the genus Chelonoidis (Testudines: Testudinidae) from the northern Bahamas, with an osteological assessment of other Neotropical tortoises". Bulletin of the Florida Museum of Natural History. 49 (1): 1–44.
- ^ Walter G. Joyce; Spencer G. Lucas; Torsten M. Scheyer; Andrew B. Heckert; Adrian P. Hunt (2009). "A thin-shelled reptile from the Late Triassic of North America and the origin of the turtle shell". Proceedings of the Royal Society B: Biological Sciences. 276 (1656): 507–513. doi:10.1098/rspb.2008.1196. PMC 2664348. PMID 18842543.
- ^ Walter G. Joyce (2017). "A Review of the Fossil Record of Basal Mesozoic Turtles" (PDF). Bulletin of the Peabody Museum of Natural History. 58 (1): 65–113. doi:10.3374/014.058.0105.
- ^ a b John Howard Hutchison (2009). "New soft-shelled turtles (Plastomeninae, Trionychidae, Testudines) from the Late Cretaceous and Paleocene of North America". PaleoBios. 29 (2): 36–47. Retrieved 7 August 2010.
- ^ a b Walter G. Joyce; Tyler R. Lyson (2011). "New Material of Gilmoremys lancensis nov. comb. (Testudines: Trionychidae) from the Hell Creek Formation and the Diagnosis of Plastomenid Turtles". Journal of Paleontology. 85 (3): 442–459. doi:10.1666/10-127.1.
- ^ Anquetin, J.; Barrett, P.M.; Jones, M.E.H.; Moore-Fay, S.; Evans, S.E. (2009). "A new stem turtle from the Middle Jurassic of Scotland: new insights into the evolution and palaeoecology of basal turtles". Proceedings of the Royal Society B. 276 (1658): 879–886. doi:10.1098/rspb.2008.1429. PMC 2664364. PMID 19019789..
- ^ Walter G. Joyce; Ariel Revan; Tyler R. Lyson; Igor G. Danilov (2009). "Two New Plastomenine Softshell Turtles from the Paleocene of Montana and Wyoming" (PDF). Bulletin of the Peabody Museum of Natural History. 50 (2): 307–325. doi:10.3374/014.050.0202. Retrieved 7 August 2010.
- ^ Gaffney, E.S.; Krause, D.W.; Zalmout, I.S. (2009). "Kinkonychelys, a new side-necked turtle (Pelomedusoides: Bothremydidae) from the Late Cretaceous of Madagascar". American Museum Novitates. 3662: 1–25. doi:10.1206/672.1. hdl:2246/5985.
- ^ Lyson, T.R.; Joyce, W.G. (2009). "A New Species of Palatobaena (Testudines: Baenidae) and a Maximum Parsimony and Bayesian Phylogenetic Analysis of Baenidae". Journal of Paleontology. 83 (3): 457–470. doi:10.1666/08-172.1.
- ^ Magdalena Borsuk−Białynicka; Susan E. Evans (2009). "A long−necked archosauromorph from the Early Triassic of Poland" (PDF). Paleontologica Polonica. 65: 203–234. Retrieved 17 August 2010.
- ^ Susan E. Evans (2009). "An early kuehneosaurid reptile (Reptilia: Diapsida) from the Early Triassic of Poland" (PDF). Paleontologica Polonica. 65: 145–178. Retrieved 13 August 2010.
- ^ Susan E. Evans; Magdalena Borsuk−Białynicka (2009). "A small lepidosauromorph reptile from the Early Triassic of Poland" (PDF). Paleontologica Polonica. 65: 179–202. Retrieved 13 August 2010.
- ^ Gasparini, Z. (2009). "A New Oxfordian Pliosaurid (Plesiosauria, Pliosauridae) in the Caribbean Seaway". Palaeontology. 52 (3): 661–669. doi:10.1111/j.1475-4983.2009.00871.x.
- ^ Druckenmiller, P.S.; Russell, A.P. (2009). "The new plesiosaurian genus Nichollssaura from Alberta, Canada: replacement name for the preoccupied genus Nichollsia". Journal of Vertebrate Paleontology. 29 (1): 276. doi:10.1080/02724634.2009.10010379.
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- ^ A.S. Schulp; N. Bardet; B. Bouya (2009). "A new species of the durophagous mosasaur Carinodens (Squamata, Mosasauridae) and additional material of Carinodens belgicus from the Maastrichtian phosphates of Morocco". Netherlands Journal of Geosciences. 88 (3): 161–167. doi:10.1017/S0016774600000871.
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