List of canids
Canidae is a family of mammals in the order Carnivora, which includes domestic dogs, wolves, coyotes, foxes, jackals, dingoes, and many other extant and extinct dog-like mammals. A member of this family is called a canid; all extant species are a part of a single subfamily, Caninae, and are called canines. They are found on all continents except Antarctica, having arrived independently or accompanied human beings over extended periods of time. Canids vary in size, including tails, from the 2 meter (6 ft 7 in) gray wolf to the 46 cm (18 in) fennec fox. Population sizes range from the Falkland Islands wolf, extinct since 1876, to the wolf, whose domestic dog subspecies has a worldwide population of over 1 billion.[1] The body forms of canids are similar, typically having long muzzles, upright ears, teeth adapted for cracking bones and slicing flesh, long legs, and bushy tails.[2] Most species are social animals, living together in family units or small groups and behaving cooperatively. Typically, only the dominant pair in a group breeds, and a litter of young is reared annually in an underground den. Canids communicate by scent signals and vocalizations.[3] One canid, the domestic dog, entered into a partnership with humans at least 14,000 years ago and today remains one of the most widely kept domestic animals.[4]
The 13 genera and 37 species of Caninae are primarily split into two tribes: Canini, which includes 9 genera and 20 species, comprising the wolf-like Canina subtribe and the South American Cerdocyonina subtribe; and Vulpini, the fox-like canids, comprising 3 genera and 14 species. Not included in either tribe is the Urocyon genus, which includes 2 species, mainly comprising the gray fox and believed to be basal to the family. In addition to the extant Caninae, Canidae comprises two extinct subfamilies designated as Hesperocyoninae and Borophaginae. Extinct species have also been placed into Caninae, in both extant and extinct genera; at least 80 extinct Caninae species have been found, as well as over 70 species in Borophaginae and nearly 30 in Hesperocyoninae, though due to ongoing research and discoveries the exact number and categorization is not fixed. The earliest canids found belong to Hesperocyoninae, and are believed to have diverged from the existing Caniformia suborder around 37 million years ago.[5]
Conventions
Conservation status | |
---|---|
EX | Extinct (2 species) |
EW | Extinct in the wild (0 species) |
CR | Critically Endangered (0 species) |
EN | Endangered (4 species) |
VU | Vulnerable (0 species) |
NT | Near threatened (5 species) |
LC | Least concern (26 species) |
Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the canid's range is provided. Ranges are based on the IUCN red list for that species unless otherwise noted. All extinct species or subspecies listed alongside extant species went extinct after 1500 CE, and are indicated by a dagger symbol "†".
Classification
The family Canidae consists of 35 extant species belonging to 12 genera and divided into 194 extant subspecies, as well the extinct genus Dusicyon, comprising two extinct species, and 13 extinct wolf subspecies, which are the only canid species to go extinct since prehistoric times. This does not include hybrid species (such as wolfdogs or coywolfs) or extinct prehistoric species (such as the dire wolf or Epicyon). Modern molecular studies indicate that the 13 genera can be grouped into 3 tribes or clades.
Subfamily Caninae
- Tribe Canini
- Subtribe Canina
- Subtribe Cerdocyonina
- Genus Atelocynus: one species
- Genus Cerdocyon: one species
- Genus Chrysocyon: one species
- Genus Dusicyon†: two species
- Genus Lycalopex: six species
- Genus Speothos: one species
- Tribe Vulpini
- Genus Nyctereutes: one species
- Genus Otocyon: one species
- Genus Vulpes: twelve species
- Genus Urocyon: two species
Canids
The following classification is based on the taxonomy described by Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis, such as the promotion of the African golden wolf to a separate species from the golden jackal. Range maps are based on IUCN range data. There are several additional proposals which are disputed, such as the promotion of the red wolf and eastern wolf as species from subspecies of the wolf, or the addition of the Italian wolf subspecies, which are not included here.
Subfamily Caninae
Tribe Canini
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population[a] |
---|---|---|---|---|
Short-eared dog | A. microtis Cabrera, 1940 Two subspecies
|
Western Amazon rainforest in South America |
Size: 72–100 cm (28–39 in) long, plus 24–35 cm (9–14 in) tail[6] Habitat: Wetlands, forest, and savanna[7] Diet: Preys primarily on fish, insects, and small mammals, as well as fruit, birds, and crabs[7][8] |
NT
|
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population[a] |
---|---|---|---|---|
African golden wolf | C. anthus[b] F. Cuvier, 1820 Six subspecies
|
North and northeastern Africa |
Size: 100 cm (39 in) long, plus 20 cm (8 in) tail[9] Habitat: Grassland, shrubland, and savanna[10] Diet: Primarily preys on wild boar and livestock, as well as other mammals and fruit[10][11] |
LC
|
Black-backed jackal | C. mesomelas Schreber, 1775 Two subspecies
|
Southern Africa and eastern Africa |
Size: 60–95 cm (24–37 in) long, plus 16–40 cm (6–16 in) tail[12] Habitat: Marine intertidal, forest, desert, grassland, shrubland, and savanna[13] Diet: Primarily preys on small to medium-sized mammals and birds[13][14] |
LC
|
Coyote | C. latrans Say, 1823 Nineteen subspecies
|
North America |
Size: 100–135 cm (39–53 in) long, plus 40 cm (16 in) tail[15] Habitat: Forest, desert, shrubland, and grassland[16] Diet: Preys on a wide variety of foods, including both small and large mammals, fruit, and insects[16] |
LC |
Ethiopian wolf | C. simensis Rüppell, 1840 Two subspecies
|
Ethiopian Highlands |
Size: 84–100 cm (33–39 in) long, plus 27–40 cm (11–16 in) tail[18] Habitat: Inland wetlands, grassland, shrubland, and rocky areas[19] Diet: Primarily preys on rodents as well as small mammals[19][20] |
EN
|
Golden jackal | C. aureus Linnaeus, 1758 Seven subspecies
|
Eastern Europe, Middle East, and southern Asia |
Size: 60–132 cm (24–52 in) long, plus 20–30 cm (8–12 in) tail[21] Habitat: Forest, grassland, shrubland, and savanna[22] Diet: Preys on a wide variety of foods, including small to large mammals, birds, fish, fruit, and insects[22][21] |
LC
|
Wolf | C. lupus Linnaeus, 1758 38 subspecies
|
Euroasia and northern North America |
Size: 105–160 cm (41–63 in) long, plus 29–50 cm (11–20 in) tail[23] Habitat: Forest, desert, rocky areas, shrubland, grassland, and inland wetlands[24] Diet: Primarily preys on large ungulates, as well as small animals, carrion, and berries[24][25] |
LC |
Side-striped jackal | C. adustus Sundevall, 1847 Seven subspecies
|
Central Africa |
Size: 69–81 cm (27–32 in) long, plus 30–41 cm (12–16 in) tail[27] Habitat: Forest, shrubland, savanna, grassland, and inland wetlands[28] Diet: Primarily preys on small to medium-sized mammals and fruit, as well as birds, insects, grass, and carrion[28][29] |
LC |
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population[a] |
---|---|---|---|---|
Crab-eating fox | C. thous Linnaeus, 1766 Five subspecies
|
Eastern and northern South America |
Size: 64 cm (25 in) long, plus 28 cm (11 in) tail[31] Habitat: Forest, savanna, shrubland, grassland, and inland wetlands[32] Diet: Primarily preys on crabs and insects, as well as rodents, birds, turtles, eggs, fruit, and carrion[31][32] |
LC
|
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population[a] |
---|---|---|---|---|
Maned wolf | C. brachyurus Illiger, 1815 |
Central South America |
Size: 100–130 cm (39–51 in) long, plus 45 cm (18 in) tail[33][34] Habitat: Forest, wetlands, grassland, shrubland, and savanna[35] Diet: Primarily preys on fruits, arthropods, and small and medium vertebrates[35] |
NT
|
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population[a] |
---|---|---|---|---|
Dhole | C. alpinus Pallas, 1811 Three subspecies
|
Southeast Asia |
Size: 90 cm (35 in) long, plus 40–45 cm (16–18 in) tail[36] Habitat: Forest, grassland, and shrubland[37] Diet: Primarily preys on ungulates, as well as small rodents and hares[37] |
EN
|
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population[a] |
---|---|---|---|---|
Falkland Islands wolf† | D. australis Kerr, 1792 |
Falkland Islands at tip of South America |
Size: Unknown Habitat: Grassland and shrubland[38] Diet: Unknown[38] |
EX |
South American fox†
|
D. avus Burmeister, 1866 |
Southern South America | Size: Unknown Habitat: Grassland and shrubland[39] Diet: Unknown[39] |
EX |
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population[a] |
---|---|---|---|---|
Culpeo | L. culpeo Molina, 1782 Six subspecies
|
Western South America |
Size: 95–132 cm (37–52 in) long, plus 32–44 cm (13–17 in) tail[41] Habitat: Forest, rocky areas, grassland, shrubland, and savanna[42] Diet: Primarily preys on rodents and lagomorphs, as well as livestock and guanacos[42][43] |
LC
|
Darwin's fox | L. fulvipes Martin, 1837 |
Limited areas in southern Chile |
Size: 48–59 cm (19–23 in) long, plus 18–26 cm (7–10 in) tail[44] Habitat: Forest and shrubland[45] Diet: Primarily preys on small mammals, insects, crabs, and fruit[44][45] |
EN
|
Hoary fox | L. vetulus Lund, 1842 |
South-central Brazil |
Size: 49–71 cm (19–28 in) long, plus 25–38 cm (10–15 in) tail[44] Habitat: Savanna[46] Diet: Primarily preys on insects, as well as small rodents, birds, reptiles, and fruit[44][46] |
LC
|
Pampas fox | L. gymnocercus Waldheim, 1814 Five subspecies
|
Southern South America |
Size: 51–74 cm (20–29 in) long, plus 25–41 cm (10–16 in) tail[44] Habitat: Forest, shrubland, and savanna[47] Diet: Primarily preys on small rodents, hares, birds, insects, and fruit, as well as carrion[44][47] |
LC
|
Sechuran fox | L. sechurae Thomas, 1900 |
Sechura Desert in southwestern Ecuador and northwestern Peru |
Size: 50–78 cm (20–31 in) long, plus 27–34 cm (11–13 in) tail[44] Habitat: Forest, desert, grassland, and shrubland[48] Diet: Primarily preys on fruit and seeds, as well as small rodents, birds, reptiles, insects, scorpions, and carrion[44][48] |
NT |
South American gray fox | L. griseus Gray, 1837 |
Southern South America |
Size: 50–66 cm (20–26 in) long, plus 12–34 cm (5–13 in) tail[44] Habitat: Forest, grassland, and shrubland[50] Diet: Primarily preys on small rodents, hares, and carrion[44][50] |
LC
|
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population[a] |
---|---|---|---|---|
African wild dog | L. pictus Temminck, 1820 Five subspecies
|
Scattered areas of Africa |
Size: 76–112 cm (30–44 in) long, plus 30–42 cm (12–17 in) tail[44] Habitat: Forest, grassland, shrubland, savanna, and desert[51] Diet: Primarily preys on medium-sized antelope[51] |
EN
|
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population[a] |
---|---|---|---|---|
Bush dog | S. venaticus Lund, 1842 Three subspecies
|
Northern South America |
Size: 57–75 cm (22–30 in) long, plus 12–15 cm (5–6 in) tail[52] Habitat: Shrubland, forest, grassland, and savanna[53] Diet: Primarily preys on small and medium mammals, as well as birds, reptiles, and fruit[53] |
NT |
Tribe Vulpini
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population[a] |
---|---|---|---|---|
Raccoon dog | N. procyonoides Gray, 1834 Five subspecies
|
Eastern Asia and Central and Eastern Europe |
Size: 49–71 cm (19–28 in) long, plus 15–23 cm (6–9 in) tail[44] Habitat: Forest, grassland, and shrubland[55] Diet: Primarily preys on insects, rodents, amphibians, birds, fish, and reptiles, as well as fruits, nuts, and berries[55] |
LC |
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population[a] |
---|---|---|---|---|
Bat-eared fox | O. megalotis Desmarest, 1822 Two subspecies
|
Southern and Eastern Africa |
Size: 46–61 cm (18–24 in) long, plus 23–34 cm (9–13 in) tail[44] Habitat: Grassland, shrubland, and savanna[57] Diet: Primarily preys on harvester termites as well as other arthropods[57] |
LC
|
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population[a] |
---|---|---|---|---|
Arctic fox | V. lagopus Linnaeus, 1758 Five subspecies
|
Arctic North America and Eurasia |
Size: 50–75 cm (20–30 in) long, plus 25–43 cm (10–17 in) tail[44] Habitat: Grassland[58] Diet: Primarily preys on lemmings, as well as other rodents, birds, and reindeer[58] |
LC
|
Bengal fox | V. bengalensis Shaw, 1800 |
India |
Size: 39–58 cm (15–23 in) long, plus 25–32 cm (10–13 in) tail[44] Habitat: Grassland and shrubland[59] Diet: Primarily preys on arthropods, rodents, reptiles, fruits, and birds[59] |
LC
|
Blanford's fox | V. cana Blanford, 1877 |
The Middle East and Central Asia |
Size: 34–47 cm (13–19 in) long, plus 26–36 cm (10–14 in) tail[44] Habitat: Desert and rocky areas[60] Diet: Primarily preys on fruit and insects[60] |
LC
|
Cape fox | V. chama A Smith, 1833 |
Southern Africa |
Size: 45–61 cm (18–24 in) long, plus 25–41 cm (10–16 in) tail[44] Habitat: Rocky areas, grassland, shrubland, and savanna[61] Diet: Primarily preys on fruit and insects[61] |
LC
|
Corsac fox | V. corsac Linnaeus, 1768 Three subspecies
|
Central Asia |
Size: 45–60 cm (18–24 in) long, plus 19–34 cm (7–13 in) tail[44] Habitat: Desert, grassland, and shrubland[62] Diet: Primarily preys on insects and small rodents[62] |
LC
|
Fennec fox | V. zerda Zimmermann, 1780 |
Northern Africa |
Size: 33–40 cm (13–16 in) long, plus 13–23 cm (5–9 in) tail[44] Habitat: Desert and marine coastal/supratidal[63] Diet: Primarily preys on rodents, insects, birds, eggs, and rabbits[63] |
LC
|
Kit fox | V. macrotis Merriam, 1888 Two subspecies
|
Western North America |
Size: 46–54 cm (18–21 in) long, plus 25–34 cm (10–13 in) tail[44] Habitat: Shrubland, savanna, and grassland[64] Diet: Primarily preys on rodents, rabbits, invertebrates, birds, lizards, and snakes[64] |
LC
|
Pale fox | V. pallida Cretzschmar, 1827 Five subspecies
|
Upper middle Africa |
Size: 38–55 cm (15–22 in) long, plus 23–29 cm (9–11 in) tail[44] Habitat: Desert, grassland, shrubland, and savanna[65] Diet: Primarily preys on plants and berries as well as rodents, reptiles, and insects[65] |
LC
|
Rüppell's fox | V. rueppellii Schinz, 1825 |
Northern Africa and the Middle East |
Size: 35–56 cm (14–22 in) long, plus 25–39 cm (10–15 in) tail[44] Habitat: Desert, shrubland, and marine coastal/supratidal[66] Diet: Primarily preys on small mammals, lizards, birds, and insects, as well as fruit and succulents[66] |
LC
|
Red fox | V. vulpes Linnaeus, 1758 44 subspecies
|
North America, Europe, Asia, and Australia |
Size: 62–72 cm (24–28 in) long, plus 40 cm (16 in) tail[67] Habitat: Shrubland, grassland, inland wetlands, forest, and desert[68] Diet: Primarily preys on small rodents, as well as birds, larger mammals, reptiles, insects, and fish[68] |
LC
|
Swift fox | V. velox Say, 1823 |
Western grasslands of North America |
Size: 48–54 cm (19–21 in) long, plus 25–34 cm (10–13 in) tail[44] Habitat: Grassland[69] Diet: Primarily preys on rabbits, mice, ground squirrels, birds, insects and lizards, as well as grasses and fruits[69] |
LC
|
Tibetan sand fox | V. ferrilata Hodgson, 1842 |
High plateaus in Nepal and western China |
Size: 49–70 cm (19–28 in) long, plus 22–29 cm (9–11 in) tail[44] Habitat: Desert, rocky areas, grassland, and shrubland[70] Diet: Primarily preys on pikas, as well as carrion and other small mammals[70] |
LC
|
Urocyon
Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population[a] |
---|---|---|---|---|
Gray fox | U. cinereoargenteus Schreber, 1775 Sixteen subspecies
|
North America and Central America |
Size: 53–66 cm (21–26 in) long, plus 28–44 cm (11–17 in) tail[44] Habitat: Forest and shrubland[71] Diet: Primarily preys on rabbits, voles, shrews, and birds, as well as insects and fruit[71] |
LC
|
Island fox | U. littoralis Baird, 1857 Six subspecies
|
Channel Islands of California |
Size: 46–63 cm (18–25 in) long, plus 12–32 cm (5–13 in) tail[44] Habitat: Marine intertidal, forest, grassland, and shrubland[72] Diet: Primarily preys on fruits, insects, birds, eggs, crabs, lizards, and small mammals[72] |
NT
|
Prehistoric canids
In addition to extant canids, a number of prehistoric species have been discovered and classified as a part of Canidae. Morphogenic and molecular phylogenic research has placed them within the extant subfamily Caninae as well as the extinct subfamilies Hesperocyoninae and Borophaginae. Within Caninae, prehistoric species have been placed into both extant genera and separate extinct genera.
The generally accepted classification of extinct canid species is primarily based for Hesperocyoninae on work by Xiaoming Wang, curator of terrestrial mammals at the Natural History Museum of Los Angeles County,[5] and on work by Wang and zoologists Richard H. Tedford and Beryl E. Taylor for Borophaginae and Caninae.[73][74][75] The species and classifications listed below are all from these works; exceptions due to more recently-described species are also listed with citations. Not all of these classifications are universally accepted, and alternate classifications for species are noted below. Where available, the approximate time period the species was extant is given in millions of years before the present (Mya), based on data from the Paleobiology Database. All listed species are extinct; where a genus, subtribe, or tribe within Caninae comprises only extinct species, it is indicated with a dagger symbol †.
Subfamily Caninae
- Tribe Canini
- Subtribe Canina
- Genus Canis
- C. antonii
- C. apolloniensis (2.6–0.78 Mya)[76]
- C. armbrusteri (Armbruster's wolf) (1.8–0.012 Mya)
- C. arnensis (Arno River dog) (2.6–0.78 Mya)
- C. brevicephalus
- C. cedazoensis (1.8–0.3 Mya)
- C. chihliensis
- C. cipio[e]
- C. dirus (Dire wolf) (1.8–0.012 Mya)
- C. edwardii (23–0.78 Mya)
- C. etruscus (Etruscan wolf) (2.6–0.13 Mya)
- C. ferox (4.9–2.6 Mya)
- C. gezi
- C. leilhardi
- C. lepophagus (4.9–0.012 Mya)
- C. longdanensis
- C. mosbachensis (Mosbach wolf) (2.6–0.13 Mya)
- C. nehringi
- C. othmani[77]
- C. palmidens
- C. variabilis
- Genus Cynotherium† (0.13–0.012 Mya)
- C. sardous (Sardinian dhole) (0.13–0.012 Mya)
- Genus Eucyon†
- E. adoxus
- E. davisi (5.3–3.6 Mya)
- E. intrepidus
- E. monticinensis
- E. odessanus
- E. skinneri (13.6–10.3 Mya)
- E. zhoui
- Genus Lycaon
- L. magnus
- L. sekowei
- Genus Mececyon†
- Genus Megacyon† (3.6–2.6 Mya)
- M. merriami (3.6–2.6 Mya)
- Genus Nurocyon†
- N. chonokhariensis
- Genus Xenocyon†[f]
- X. africanus
- X. antonii
- X. falconeri
- X. lycaonoides (0.78–0.3 Mya)
- Genus Canis
- Subtribe Cerdocyonina
- Genus Cerdocyon
- Genus Nyctereutes
- N. abdeslami (3.6–2.6 Mya)
- N. donnezani
- N. megamastoides (2.6–0.13 Mya)
- N. sinensis
- N. terblanchei
- N. tingi
- Genus Protocyon†
- P. orcesi (0.78–0.012 Mya)
- P. scagliarum
- P. troglodytes (0.13–0.012 Mya)
- Genus Speothos
- S. pacivorus (Pleistocene bush dog)
- Genus Theriodictis† (1.8 Mya)
- T. floridanus (1.8 Mya)
- Subtribe Canina
- Tribe Vulpini
- Genus Metalopex† (10–4.9 Mya)
- M. bakeri (10–4.9 Mya)
- M. macconnelli (10–5.3 Mya)
- M. merriami (10–5.3 Mya)
- Genus Prototocyon
- P. curvipalatus
- P. recki
- Genus Vulpes
- V. alopecoides (2.5–0.13 Mya)
- V. angustidens
- V. beihaiensis
- V. chikushanensis
- V. galaticus
- V. praecorsac (3.2–0.78 Mya)
- V. praeglacialis
- V. riffautae
- V. skinneri
- V. stenognathus (14–0.3 Mya)
- V. qiuzhudingi[79]
- Genus Metalopex† (10–4.9 Mya)
- Urocyon
- Genus Urocyon
- U. minicephalus (1.8–0.3 Mya)
- U. progressus (4.9–1.8 Mya)
- Genus Urocyon
- Basal Caninae
- Genus Leptocyon† (31–10 Mya)
- L. delicatus (31–20 Mya)
- L. douglassi (31–26 Mya)
- L. gregorii (25–20 Mya)
- L. leidyi (20–14 Mya)
- L. matthewi (14–10 Mya)
- L. mollis (31–20 Mya)
- L. tejonensis (14–10 Mya)
- L. vafer (14–10 Mya)
- L. vulpinus (20–16 Mya)
- Genus Leptocyon† (31–10 Mya)
- Unclassified
- Genus Protemnocyon† (34–33 Mya)
- P. inflatus (34–33 Mya)
- Genus Protemnocyon† (34–33 Mya)
Subfamily Borophaginae
- Tribe Borophagini (26–1.8 Mya)
- Genus Cormocyon (26–20 Mya)
- C. copei (26–20 Mya)
- C. haydeni (25–20 Mya)
- Genus Desmocyon (20–16 Mya)
- D. matthewi (20–16 Mya)
- D. thomsoni (20–16 Mya)
- Genus Euoplocyon (20–14 Mya)
- E. brachygnathus (16–14 Mya)
- E. spissidens (20–16 Mya)
- Genus Metatomarctus (20–16 Mya)
- M. canavus (20–16 Mya)
- Genus Microtomarctus (16–14 Mya)
- M. conferta (16–14 Mya)
- Genus Protomarctus (16–14 Mya)
- P. optatus (16–14 Mya)
- Genus Psalidocyon (16–14 Mya)
- P. marianae (16–14 Mya)
- Genus Tephrocyon (16–14 Mya)
- T. rurestris (16–14 Mya)
- Subtribe Aelurodontina (16–5.3 Mya)
- Subtribe Borophagina (16–1.8 Mya)
- Genus Borophagus (14–1.8 Mya)
- B. diversidens (4.9–1.8 Mya)
- B. dudleyi (5.3–3.6 Mya)
- B. hilli (5.3–3.6 Mya)
- B. littoralis (14–10 Mya)
- B. orc (5.3–4.9 Mya)
- B. parvus (10–4.9 Mya)
- B. pugnator (10–5.3 Mya)
- B. secundus (10–4.9 Mya)
- Genus Carpocyon (16–5.3 Mya)
- C. compressus (16–14 Mya)
- C. limosus (10–5.3 Mya)
- C. robustus (14–10 Mya)
- C. webbi (14–5.3 Mya)
- Genus Epicyon (16–4.9 Mya)
- E. aelurodontoides (10–4.9 Mya)
- E. haydeni (10–4.9 Mya)
- E. saevus (16–4.9 Mya)
- Genus Paratomarctus (16–5.3 Mya)
- P. euthos (14–10 Mya)
- P. temerarius (16–5.3 Mya)
- Genus Protepicyon (16–14 Mya)
- P. raki (16–14 Mya)
- Genus Borophagus (14–1.8 Mya)
- Subtribe Cynarctina (16–10 Mya)
- Genus Cynarctus (16–10 Mya)
- C. crucidens (12–10 Mya)
- C. galushai (16–14 Mya)
- C. marylandica (16–14 Mya)
- C. saxatilis (16–14 Mya)
- C. voorhiesi (14–10 Mya)
- C. wangi (16–14 Mya)[81]
- Genus Paracynarctus (16–14 Mya)
- P. kelloggi (16–14 Mya)
- P. sinclairi (16–14 Mya)
- Genus Cynarctus (16–10 Mya)
- Genus Cormocyon (26–20 Mya)
- Tribe Phlaocyonini (30.8–13.6 Mya)
- Genus Cynarctoides (31–14 Mya)
- C. acridens (20–14 Mya)
- C. emryi (20–16 Mya)
- C. gawnae (20–16 Mya)
- C. harlowi (25–20 Mya)
- C. lemur (31–20 Mya)
- C. luskensis (25–20 Mya)
- C. roii (31–26 Mya)
- Genus Phlaocyon (31–16 Mya)
- P. achoros (25—20 Mya)
- P. annectens (25–20 Mya)
- P. latidens (31–20 Mya)
- P. leucosteus (20–16 Mya)
- P. mariae(20–16 Mya)
- P. marslandensis (20–16 Mya)
- P. minor (25–16 Mya)
- P. multicuspus (25–20 Mya)
- P. taylori (31–25 Mya)[82]
- P. yatkolai (20–16 Mya)
- Genus Cynarctoides (31–14 Mya)
- Basal Borophaginae
- Genus Archaeocyon (31–20 Mya)
- A. falkenbachi (31–20 Mya)
- A. leptodus (31–26 Mya)
- A. pavidus (31–26 Mya)
- Genus Otarocyon (34–26 Mya)
- O. cooki (31–26 Mya)
- O. macdonaldi (34–33 Mya)
- Genus Oxetocyon (33–31 Mya)
- O. cuspidatus (33–31 Mya)
- Genus Rhizocyon (31–20 Mya)
- R. oregonensis (31–20 Mya)
- Genus Archaeocyon (31–20 Mya)
Subfamily Hesperocyoninae
- Genus Cynodesmus (31–20 Mya)
- C. martini (31–20 Mya)
- C. thooides (31–26 Mya)
- Genus Caedocyon (31–20 Mya)
- C. tedfordi (31–20 Mya)
- Genus Ectopocynus (31–16 Mya)
- E. antiquus (31–20 Mya)
- E. intermedius (31–20 Mya)
- E. siplicidens (20–16 Mya)
- Genus Enhydrocyon (31–20 Mya)
- E. basilatus (25–20 Mya)
- E. crassidens (26–20 Mya)
- E. pahinsintewkpa (26–20 Mya)
- E. stenocephalus (31–20 Mya)
- E. sectorius
- Genus Hesperocyon (37–31 Mya)
- H. coloradensis (34–33 Mya)
- H. gregarius (37–31 Mya)
- Genus Mesocyon (33–20 Mya)
- M. brachyops (31–20 Mya)
- M. coryphaeus (31–20 Mya)
- M. temnodon (33–20 Mya)
- Genus Osbornodon (33–14 Mya)
- O. brachypus (20–16 Mya)
- O. fricki (16–14 Mya)
- O. iamonensis (20–16 Mya)
- O. renjiei (33–31 Mya)
- O. scitulus (21–16 Mya)[83]
- O. sesnoni (31–20 Mya)
- O. wangi (31–20 Mya)[82]
- Genus Paraenhydrocyon (25–20 Mya)
- P. josephi (25–20 Mya)
- P. robustus (25–20 Mya)
- P. wallovianus (25–20 Mya)
- Genus Philotrox (31–26 Mya)
- P. condoni (31–26 Mya)
- Genus Prohesperocyon (37–34 Mya)
- P. wilsoni (37–34 Mya)
- Genus Sunkahetanka (31–26 Mya)
- S. geringensis (31–26 Mya)
Notes
- ^ a b c d e f g h i j k l m Population figures rounded to the nearest hundred. Population trends as described by the International Union for Conservation of Nature.
- ^ It has been proposed that the binomial for the African golden wolf be changed to Canis lupaster, which the IUCN has adopted but is not yet universal
- ^ The Falklands Island wolf is believed to have been driven extinct in 1876[38]
- ^ The South American fox is believed to have gone extinct sometime between 1454 and 1626[40]
- ^ Also potentially placed in the Eucyon genus
- ^ Xenocyon is sometimes considered a subgenus of Canis
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