Wheat (Triticum spp.) is a cereal grain, (botanically, a type of fruit called a caryopsis) originally from the Levant region of the Near East but now cultivated worldwide. In 2013, world production of wheat was 713 million tons, making it the third most-produced cereal after maize (1,016 million tons) and rice (745 million tons). Wheat was the second most-produced cereal in 2009; world production in that year was 682 million tons, after maize (817 million tons), and with rice as a close third (679 million tons).
This grain is grown on more land area than any other commercial food. World trade in wheat is greater than for all other crops combined. Globally, wheat is the leading source of vegetable protein in human food, having a higher protein content than other major cereals, maize (corn), or rice. In terms of total production tonnages used for food, it is currently second to rice as the main human food crop and ahead of maize, after allowing for maize's more extensive use in animal feeds. The archaeological record suggests that this first occurred in the regions known as the Fertile Crescent.
- 1 Origin
- 2 Farming techniques
- 3 Genetics
- 4 Plant breeding
- 5 Hulled versus free-threshing wheat
- 6 Naming
- 7 As a food
- 8 Commercial use
- 9 Production and consumption
- 10 Agronomy
- 11 Diseases
- 12 See also
- 13 References
- 14 Further reading
- 15 External links
Cultivation and repeated harvesting and sowing of the grains of wild grasses led to the creation of domestic strains, as mutant forms ('sports') of wheat were preferentially chosen by farmers. In domesticated wheat, grains are larger, and the seeds (inside the spikelets) remain attached to the ear by a toughened rachis during harvesting. In wild strains, a more fragile rachis allows the ear to easily shatter and disperse the spikelets. Selection for these traits by farmers might not have been deliberately intended, but simply have occurred because these traits made gathering the seeds easier; nevertheless such 'incidental' selection was an important part of crop domestication. As the traits that improve wheat as a food source also involve the loss of the plant's natural seed dispersal mechanisms, highly domesticated strains of wheat cannot survive in the wild.
Cultivation of wheat began to spread beyond the Fertile Crescent after about 8000 BCE. Jared Diamond traces the spread of cultivated emmer wheat starting in the Fertile Crescent sometime before 8800 BCE. Archaeological analysis of wild emmer indicates that it was first cultivated in the southern Levant with finds dating back as far as 9600 BCE. Genetic analysis of wild einkorn wheat suggests that it was first grown in the Karacadag Mountains in southeastern Turkey. Dated archeological remains of einkorn wheat in settlement sites near this region, including those at Abu Hureyra in Syria, suggest the domestication of einkorn near the Karacadag Mountain Range. With the anomalous exception of two grains from Iraq ed-Dubb, the earliest carbon-14 date for einkorn wheat remains at Abu Hureyra is 7800 to 7500 years BCE.
Remains of harvested emmer from several sites near the Karacadag Range have been dated to between 8600 (at Cayonu) and 8400 BCE (Abu Hureyra), that is, in the Neolithic period. With the exception of Iraq ed-Dubb, the earliest carbon-14 dated remains of domesticated emmer wheat were found in the earliest levels of Tell Aswad, in the Damascus basin, near Mount Hermon in Syria. These remains were dated by Willem van Zeist and his assistant Johanna Bakker-Heeres to 8800 BCE. They also concluded that the settlers of Tell Aswad did not develop this form of emmer themselves, but brought the domesticated grains with them from an as yet unidentified location elsewhere.
The cultivation of emmer reached Greece, Cyprus and India by 6500 BCE, Egypt shortly after 6000 BCE, and Germany and Spain by 5000 BCE. "The early Egyptians were developers of bread and the use of the oven and developed baking into one of the first large-scale food production industries."  By 3000 BCE, wheat had reached the British Isles and Scandinavia. A millennium later it reached China. The first identifiable bread wheat (Triticum aestivum) with sufficient gluten for yeasted breads has been identified using DNA analysis in samples from a granary dating to approximately 1350 BCE at Assiros in Greek Macedonia.
From Asia, wheat continued to spread throughout Europe. In the British Isles, wheat straw (thatch) was used for roofing in the Bronze Age, and was in common use until the late 19th century. 
Technological advances in soil preparation and seed placement at planting time, use of crop rotation and fertilizers to improve plant growth, and advances in harvesting methods have all combined to promote wheat as a viable crop. Agricultural cultivation using horse collar leveraged plows (at about 3000 BCE) was one of the first innovations that increased productivity. Much later, when the use of seed drills replaced broadcasting sowing of seed in the 18th century, another great increase in productivity occurred.
Yields of pure wheat per unit area increased as methods of crop rotation were applied to long cultivated land, and the use of fertilizers became widespread. Improved agricultural husbandry has more recently included threshing machines and reaping machines (the 'combine harvester'), tractor-drawn cultivators and planters, and better varieties (see Green Revolution and Norin 10 wheat). Great expansion of wheat production occurred as new arable land was farmed in the Americas and Australia in the 19th and 20th centuries.
Wheat genetics is more complicated than that of most other domesticated species. Some wheat species are diploid, with two sets of chromosomes, but many are stable polyploids, with four sets of chromosomes (tetraploid) or six (hexaploid).
- Einkorn wheat (T. monococcum) is diploid (AA, two complements of seven chromosomes, 2n=14).
- Most tetraploid wheats (e.g. emmer and durum wheat) are derived from wild emmer, T. dicoccoides. Wild emmer is itself the result of a hybridization between two diploid wild grasses, T. urartu and a wild goatgrass such as Aegilops searsii or Ae. speltoides. The unknown grass has never been identified among now surviving wild grasses, but the closest living relative is Aegilops speltoides. The hybridization that formed wild emmer (AABB) occurred in the wild, long before domestication, and was driven by natural selection.
- Hexaploid wheats evolved in farmers' fields. Either domesticated emmer or durum wheat hybridized with yet another wild diploid grass (Aegilops tauschii) to make the hexaploid wheats, spelt wheat and bread wheat. These have three sets of paired chromosomes, three times as many as in diploid wheat.
The presence of certain versions of wheat genes has been important for crop yields. Apart from mutant versions of genes selected in antiquity during domestication, there has been more recent deliberate selection of alleles that affect growth characteristics. Genes for the 'dwarfing' trait, first used by Japanese wheat breeders to produce short-stalked wheat, have had a huge effect on wheat yields world-wide, and were major factors in the success of the Green Revolution in Mexico and Asia, an initiative led by Norman Borlaug. Dwarfing genes enable the carbon that is fixed in the plant during photosynthesis to be diverted towards seed production, and they also help prevent the problem of lodging. 'Lodging' occurs when an ear stalk falls over in the wind and rots on the ground, and heavy nitrogenous fertilization of wheat makes the grass grow taller and become more susceptible to this problem. By 1997, 81% of the developing world's wheat area was planted to semi-dwarf wheats, giving both increased yields and better response to nitrogenous fertilizer.
Heterosis, or hybrid vigor (as in the familiar F1 hybrids of maize), occurs in common (hexaploid) wheat, but it is difficult to produce seed of hybrid cultivars on a commercial scale (as is done with maize) because wheat flowers are perfect and normally self-pollinate. Commercial hybrid wheat seed has been produced using chemical hybridizing agents; these chemicals selectively interfere with pollen development, or naturally occurring cytoplasmic male sterility systems. Hybrid wheat has been a limited commercial success in Europe (particularly France), the United States and South Africa. F1 hybrid wheat cultivars should not be confused with the standard method of breeding inbred wheat cultivars by crossing two lines using hand emasculation, then selfing or inbreeding the progeny many (ten or more) generations before release selections are identified to be released as a variety or cultivar.
Synthetic hexaploids made by crossing the wild goatgrass wheat ancestor Aegilops tauschii and various durum wheats are now being deployed, and these increase the genetic diversity of cultivated wheats.
Stomata (or leaf pores) are involved in both uptake of carbon dioxide gas from the atmosphere and water vapor losses from the leaf due to water transpiration. Basic physiological investigation of these gas exchange processes has yielded valuable carbon isotope based methods that are used for breeding wheat varieties with improved water-use efficiency. These varieties can improve crop productivity in rain-fed dry-land wheat farms.
In 2010, a team of UK scientists funded by BBSRC announced they had decoded the wheat genome for the first time (95% of the genome of a variety of wheat known as Chinese Spring line 42). This genome was released in a basic format for scientists and plant breeders to use but was not a fully annotated sequence which was reported in some of the media.
On 29 November 2012, an essentially complete gene set of bread wheat has been published. Random shotgun libraries of total DNA and cDNA from the T. aestivum cv. Chinese Spring (CS42) were sequenced in Roche 454 pyrosequencer using GS FLX Titanium and GS FLX+ platforms to generate 85 Gb of sequence (220 million reads), equivalent to 5X genome coverage and identified between 94,000 and 96,000 genes.
This sequence data provides direct access to about 96,000 genes, relying on orthologous gene sets from other cereals. and represents an essential step towards a systematic understanding of biology and engineering the cereal crop for valuable traits. Its implications in cereal genetics and breeding includes the examination of genome variation, association mapping using natural populations, performing wide crosses and alien introgression, studying the expression and nucleotide polymorphism in transcriptomes, analyzing population genetics and evolutionary biology, and studying the epigenetic modifications. Moreover, the availability of large-scale genetic markers generated through NGS technology will facilitate trait mapping and make marker-assisted breeding much feasible.
Moreover, the data not only facilitate in deciphering the complex phenomena such as heterosis and epigenetics, it may also enable breeders to predict which fragment of a chromosome is derived from which parent in the progeny line, thereby recognizing crossover events occurring in every progeny line and inserting markers on genetic and physical maps without ambiguity. In due course, this will assist in introducing specific chromosomal segments from one cultivar to another. Besides, the researchers had identified diverse classes of genes participating in energy production, metabolism and growth that were probably linked with crop yield, which can now be utilized for the development of transgenic wheat. Thus whole genome sequence of wheat and the availability of thousands of SNPs will inevitably permit the breeders to stride towards identifying novel traits, providing biological knowledge and empowering biodiversity-based breeding.
In traditional agricultural systems wheat populations often consist of landraces, informal farmer-maintained populations that often maintain high levels of morphological diversity. Although landraces of wheat are no longer grown in Europe and North America, they continue to be important elsewhere. The origins of formal wheat breeding lie in the nineteenth century, when single line varieties were created through selection of seed from a single plant noted to have desired properties. Modern wheat breeding developed in the first years of the twentieth century and was closely linked to the development of Mendelian genetics. The standard method of breeding inbred wheat cultivars is by crossing two lines using hand emasculation, then selfing or inbreeding the progeny. Selections are identified (shown to have the genes responsible for the varietal differences) ten or more generations before release as a variety or cultivar.
The major breeding objectives include high grain yield, good quality, disease and insect resistance and tolerance to abiotic stresses, including mineral, moisture and heat tolerance. The major diseases in temperate environments include the following, arranged in a rough order of their significance from cooler to warmer climates: eyespot, Stagonospora nodorum blotch (also known as glume blotch), yellow or stripe rust, powdery mildew, Septoria tritici blotch (sometimes known as leaf blotch), brown or leaf rust, Fusarium head blight, tan spot and stem rust. In tropical areas, spot blotch (also known as Helminthosporium leaf blight) is also important.
Wheat has also been the subject of mutation breeding, with the use of gamma, x-rays, ultraviolet light, and sometimes harsh chemicals. The varieties of wheat created through these methods are in the hundreds (going as far back as 1960), more of them being created in higher populated countries such as China. Bread wheat with high grain iron and zinc content was developed through gamma radiation breeding.
Because wheat self-pollinates, creating hybrid varieties is extremely labor-intensive; the high cost of hybrid wheat seed relative to its moderate benefits have kept farmers from adopting them widely despite nearly 90 years of effort. F1 hybrid wheat cultivars should not be confused with wheat cultivars deriving from standard plant breeding. Heterosis or hybrid vigor (as in the familiar F1 hybrids of maize) occurs in common (hexaploid) wheat, but it is difficult to produce seed of hybrid cultivars on a commercial scale as is done with maize because wheat flowers are perfect in the botanical sense, meaning they have both male and female parts, and normally self-pollinate. Commercial hybrid wheat seed has been produced using chemical hybridizing agents, plant growth regulators that selectively interfere with pollen development, or naturally occurring cytoplasmic male sterility systems. Hybrid wheat has been a limited commercial success in Europe (particularly France), the United States and South Africa.
Hulled versus free-threshing wheat
The four wild species of wheat, along with the domesticated varieties einkorn, emmer and spelt, have hulls. This more primitive morphology (in evolutionary terms) consists of toughened glumes that tightly enclose the grains, and (in domesticated wheats) a semi-brittle rachis that breaks easily on threshing. The result is that when threshed, the wheat ear breaks up into spikelets. To obtain the grain, further processing, such as milling or pounding, is needed to remove the hulls or husks. In contrast, in free-threshing (or naked) forms such as durum wheat and common wheat, the glumes are fragile and the rachis tough. On threshing, the chaff breaks up, releasing the grains. Hulled wheats are often stored as spikelets because the toughened glumes give good protection against pests of stored grain.
There are many botanical classification systems used for wheat species, discussed in a separate article on wheat taxonomy. The name of a wheat species from one information source may not be the name of a wheat species in another.
Within a species, wheat cultivars are further classified by wheat breeders and farmers in terms of:
- Growing season, such as winter wheat vs. spring wheat.
- Protein content. Bread wheat protein content ranges from 10% in some soft wheats with high starch contents, to 15% in hard wheats.
- The quality of the wheat protein gluten. This protein can determine the suitability of a wheat to a particular dish. A strong and elastic gluten present in bread wheats enables dough to trap carbon dioxide during leavening, but elastic gluten interferes with the rolling of pasta into thin sheets. The gluten protein in durum wheats used for pasta is strong but not elastic.
- Grain color (red, white or amber). Many wheat varieties are reddish-brown due to phenolic compounds present in the bran layer which are transformed to pigments by browning enzymes. White wheats have a lower content of phenolics and browning enzymes, and are generally less astringent in taste than red wheats. The yellowish color of durum wheat and semolina flour made from it is due to a carotenoid pigment called lutein, which can be oxidized to a colorless form by enzymes present in the grain.
Major cultivated species of wheat
- Common wheat or bread wheat (T. aestivum) – A hexaploid species that is the most widely cultivated in the world.
- Spelt (T. spelta) – Another hexaploid species cultivated in limited quantities. Spelt is sometimes considered a subspecies of the closely related species common wheat (T. aestivum), in which case its botanical name is considered to be Triticum aestivum subsp. spelta.
- Durum (T. durum) – The only tetraploid form of wheat widely used today, and the second most widely cultivated wheat.
- Emmer (T. dicoccon) – A tetraploid species, cultivated in ancient times but no longer in widespread use.
- Khorasan (Triticum turgidum ssp. turanicum also called Triticum turanicum) is a tetraploid wheat species. It is an ancient grain type; Khorasan refers to a historical region in modern-day Afghanistan and the northeast of Iran. This grain is twice the size of modern-day wheat and is known for its rich nutty flavor.
- Einkorn (T. monococcum) – A diploid species with wild and cultivated variants. Domesticated at the same time as emmer wheat, but never reached the same importance.
Classes used in the United States:
- Durum – Very hard, translucent, light-colored grain used to make semolina flour for pasta & bulghur; high in protein, specifically, gluten protein.
- Hard Red Spring – Hard, brownish, high-protein wheat used for bread and hard baked goods. Bread Flour and high-gluten flours are commonly made from hard red spring wheat. It is primarily traded at the Minneapolis Grain Exchange.
- Hard Red Winter – Hard, brownish, mellow high-protein wheat used for bread, hard baked goods and as an adjunct in other flours to increase protein in pastry flour for pie crusts. Some brands of unbleached all-purpose flours are commonly made from hard red winter wheat alone. It is primarily traded on the Kansas City Board of Trade. One variety is known as "turkey red wheat", and was brought to Kansas by Mennonite immigrants from Russia.
- Soft Red Winter – Soft, low-protein wheat used for cakes, pie crusts, biscuits, and muffins. Cake flour, pastry flour, and some self-rising flours with baking powder and salt added, for example, are made from soft red winter wheat. It is primarily traded on the Chicago Board of Trade.
- Hard White – Hard, light-colored, opaque, chalky, medium-protein wheat planted in dry, temperate areas. Used for bread and brewing.
- Soft White – Soft, light-colored, very low protein wheat grown in temperate moist areas. Used for pie crusts and pastry. Pastry flour, for example, is sometimes made from soft white winter wheat.
Red wheats may need bleaching; therefore, white wheats usually command higher prices than red wheats on the commodities market.
As a food
|Nutritional value per 100 g (3.5 oz)|
|Energy||1,368 kJ (327 kcal)|
|Dietary fiber||12.2 g|
|Pantothenic acid (B5)||
|Percentages are roughly approximated using US recommendations for adults.
Source: USDA Nutrient Database
Raw wheat can be ground into flour or, using hard durum wheat only, can be ground into semolina; germinated and dried creating malt; crushed or cut into cracked wheat; parboiled (or steamed), dried, crushed and de-branned into bulgur also known as groats. If the raw wheat is broken into parts at the mill, as is usually done, the outer husk or bran can be used several ways. Wheat is a major ingredient in such foods as bread, porridge, crackers, biscuits, Muesli, pancakes, pies, pastries, cakes, cookies, muffins, rolls, doughnuts, gravy, boza (a fermented beverage), and breakfast cereals (e.g., Wheatena, Cream of Wheat, Shredded Wheat, and Wheaties).
In 100 grams, wheat provides 327 calories and is an excellent source (more than 19% of the Daily Value, DV) of multiple essential nutrients, such as protein, dietary fiber, manganese, phosphorus and niacin (table). Several B vitamins and other dietary minerals are in significant content (table). Wheat is 13% water, 71% carbohydrates, 1.5% fat and 13% protein (table).
|cooking Reduction %||10||30||20||25||25||35||0||0||30||10||15||20||10||20||5||10||25|
Ch. = Choline; Ca = Calcium; Fe = Iron; Mg = Magnesium; P = Phosphorus; K = Potassium; Na = Sodium; Zn = Zinc; Cu = Copper; Mn = Manganese; Se = Selenium; %DV = % daily value i.e. % of DRI (Dietary Reference Intake) Note: All nutrient values including protein are in %DV per 100 grams of the food item. Significant values are highlighted in light Gray color and bold letters. Cooking reduction = % Maximum typical reduction in nutrients due to boiling without draining for ovo-lacto-vegetables group Q = Quality of Protein in terms of completeness without adjusting for digestability.
100 g (3.5 oz) of hard red winter wheat contain about 12.6 g (0.44 oz) of protein, 1.5 g (0.053 oz) of total fat, 71 g (2.5 oz) of carbohydrate (by difference), 12.2 g (0.43 oz) of dietary fiber, and 3.2 mg (0.00011 oz) of iron (17% of the daily requirement); the same weight of hard red spring wheat contains about 15.4 g (0.54 oz) of protein, 1.9 g (0.067 oz) of total fat, 68 g (2.4 oz) of carbohydrate (by difference), 12.2 g (0.43 oz) of dietary fiber, and 3.6 mg (0.00013 oz) of iron (20% of the daily requirement).
Wheat is grown on more than 218,000,000 hectares (540,000,000 acres), larger than for any other crop. World trade in wheat is greater than for all other crops combined. With rice, wheat is the world's most favored staple food. It is a major diet component because of the wheat plant's agronomic adaptability with the ability to grow from near arctic regions to equator, from sea level to plains of Tibet, approximately 4,000 m (13,000 ft) above sea level. In addition to agronomic adaptability, wheat offers ease of grain storage and ease of converting grain into flour for making edible, palatable, interesting and satisfying foods. Wheat is the most important source of carbohydrate in a majority of countries.
Wheat protein is easily digested by nearly 99% of the human population (all but those with gluten-related disorders), as is its starch. With a small amount of animal or legume protein added, a wheat-based meal is highly nutritious.
The most common forms of wheat are white and red wheat. However, other natural forms of wheat exist. Other commercially minor but nutritionally promising species of naturally evolved wheat species include black, yellow and blue wheat.
While coeliac disease is caused by a reaction to wheat proteins, it is not the same as a wheat allergy. Other diseases triggered by gluten consumption are non-celiac gluten sensitivity, (estimated in one study to affect the general population in a wide range from 0.5% to 13%), gluten ataxia and dermatitis herpetiformis.
Comparison of wheat with other major staple foods
The following table shows the nutrient content of wheat and other major staple foods in a raw form.
Raw forms of these staples, however, are not edible and cannot be digested. These must be sprouted, or prepared and cooked as appropriate for human consumption. In sprouted or cooked form, the relative nutritional and anti-nutritional contents of each of these grains is remarkably different from that of raw form of these grains reported in this table.
In cooked form, the nutrition value for each staple depends on the cooking method (for example: baking, boiling, steaming, frying, etc.).
|STAPLE:||RDA||Maize / Corn[A]||Rice (white)[B]||Rice (brown)[I]||Wheat[C]||Potato[D]||Cassava[E]||Soybean (Green)[F]||Sweet potato[G]||Sorghum[H]||Yam[Y]||Plantain[Z]|
|Component (per 100g portion)||Amount||Amount||Amount||Amount||Amount||Amount||Amount||Amount||Amount||Amount||Amount||Amount|
|Vitamin C (mg)||90||0||0||0||0||19.7||20.6||29||2.4||0||17.1||18.4|
|Niacin (B3) (mg)||16||3.63||1.6||5.09||5.46||1.05||0.85||1.65||0.56||2.93||0.55||0.69|
|Pantothenic acid (B5) (mg)||5||0.42||1.01||1.49||0.95||0.30||0.11||0.15||0.80||-||0.31||0.26|
|Vitamin B6 (mg)||1.3||0.62||0.16||0.51||0.3||0.30||0.09||0.07||0.21||-||0.29||0.30|
|Folate Total (B9) (μg)||400||19||8||20||38||16||27||165||11||0||23||22|
|Vitamin A (IU)||5000||214||0||0||9||2||13||180||14187||0||138||1127|
|Vitamin E, alpha-tocopherol (mg)||15||0.49||0.11||0.59||1.01||0.01||0.19||0||0.26||0||0.39||0.14|
|Vitamin K1 (μg)||120||0.3||0.1||1.9||1.9||1.9||1.9||0||1.8||0||2.6||0.7|
|Saturated fatty acids (g)||0.67||0.18||0.58||0.26||0.03||0.07||0.79||0.02||0.46||0.04||0.14|
|Monounsaturated fatty acids (g)||1.25||0.21||1.05||0.2||0.00||0.08||1.28||0.00||0.99||0.01||0.03|
|Polyunsaturated fatty acids (g)||2.16||0.18||1.04||0.63||0.04||0.05||3.20||0.01||1.37||0.08||0.07|
|A corn, yellow||B rice, white, long-grain, regular, raw, unenriched|
|C wheat, hard red winter||D potato, flesh and skin, raw|
|E cassava, raw||F soybeans, green, raw|
|G sweet potato, raw, unprepared||H sorghum, raw|
|Y yam, raw||Z plantains, raw|
|I rice, brown, long-grain, raw|
Harvested wheat grain that enters trade is classified according to grain properties for the purposes of the commodity markets. Wheat buyers use these to decide which wheat to buy, as each class has special uses, and producers use them to decide which classes of wheat will be most profitable to cultivate.
Wheat is widely cultivated as a cash crop because it produces a good yield per unit area, grows well in a temperate climate even with a moderately short growing season, and yields a versatile, high-quality flour that is widely used in baking. Most breads are made with wheat flour, including many breads named for the other grains they contain, for example, most rye and oat breads. The popularity of foods made from wheat flour creates a large demand for the grain, even in economies with significant food surpluses.
In recent years, low international wheat prices have often encouraged farmers in the United States to change to more profitable crops. In 1998, the price at harvest was $2.68 per bushel. USDA report revealed that in 1998, average operating costs were $1.43 per bushel and total costs were $3.97 per bushel. In that study, farm wheat yields averaged 41.7 bushels per acre (2.2435 metric ton/hectare), and typical total wheat production value was $31,900 per farm, with total farm production value (including other crops) of $173,681 per farm, plus $17,402 in government payments. There were significant profitability differences between low- and high-cost farms, mainly due to crop yield differences, location, and farm size.
In 2007 there was a dramatic rise in the price of wheat due to freezes and flooding in the Northern Hemisphere and a drought in Australia. Wheat futures in September, 2007 for December and March delivery had risen above $9.00 a bushel, prices never seen before. There were complaints in Italy about the high price of pasta.
Production and consumption
In 2011, global per capita wheat consumption was 65 kg (143 lb), with the highest per capita consumption of 210 kg (460 lb) found in Azerbaijan. In 1997, global wheat consumption was 101 kg (223 lb) per capita, with the highest consumption 623 kg (1,373 lb) per capita in Denmark, but most of this (81%) was for animal feed. Wheat is the primary food staple in North Africa and the Middle East, and is growing in popularity in Asia. Unlike rice, wheat production is more widespread globally though China's share is almost one-sixth of the world.
"There is a little increase in yearly crop yield comparison to the year 1990. The reason for this is not in development of sowing area, but the slow and successive increasing of the average yield. Average 2.5 tons wheat was produced on one hectare crop land in the world in the first half of 1990s, however this value was about 3 tons in 2009. In the world per capita wheat producing area continuously decreased between 1990 and 2009 considering the change of world population. There was no significant change in wheat producing area in this period. However, due to the improvement of average yields there is some fluctuation in each year considering the per capita production, but there is no considerable decline. In 1990 per capita production was 111.98 kg/capita/year, while it was already 100.62 kg/capita/year in 2009. The decline is evident and the per capita production level of the year 1990 can not be feasible simultaneously with the growth of world population in spite of the increased average yields. In the whole period the lowest per capita production was in 2006."
In the 20th century, global wheat output expanded by about 5-fold, but until about 1955 most of this reflected increases in wheat crop area, with lesser (about 20%) increases in crop yields per unit area. After 1955 however, there was a ten-fold increase in the rate of wheat yield improvement per year, and this became the major factor allowing global wheat production to increase. Thus technological innovation and scientific crop management with synthetic nitrogen fertilizer, irrigation and wheat breeding were the main drivers of wheat output growth in the second half of the century. There were some significant decreases in wheat crop area, for instance in North America.
Better seed storage and germination ability (and hence a smaller requirement to retain harvested crop for next year's seed) is another 20th century technological innovation. In Medieval England, farmers saved one-quarter of their wheat harvest as seed for the next crop, leaving only three-quarters for food and feed consumption. By 1999, the global average seed use of wheat was about 6% of output.
Several factors are currently slowing the rate of global expansion of wheat production: population growth rates are falling while wheat yields continue to rise, and the better economic profitability of other crops such as soybeans and maize, linked with investment in modern genetic technologies, has promoted shifts to other crops.
In the Punjab region of India and Pakistan, as well as North China, irrigation has been a major contributor to increased grain output. More widely over the last 40 years, a massive increase in fertilizer use together with the increased availability of semi-dwarf varieties in developing countries, has greatly increased yields per hectare. In developing countries, use of (mainly nitrogenous) fertilizer increased 25-fold in this period. However, farming systems rely on much more than fertilizer and breeding to improve productivity. A good illustration of this is Australian wheat growing in the southern winter cropping zone, where, despite low rainfall (300 mm), wheat cropping is successful even with relatively little use of nitrogenous fertilizer. This is achieved by 'rotation cropping' (traditionally called the ley system) with leguminous pastures and, in the last decade, including a canola crop in the rotations has boosted wheat yields by a further 25%. In these low rainfall areas, better use of available soil-water (and better control of soil erosion) is achieved by retaining the stubble after harvesting and by minimizing tillage.
In 2009, the most productive farms for wheat were in France producing 7.45 metric tonnes per hectare (although French production has low protein content and requires blending with higher protein wheat to meet the specifications required in some countries). The five largest producers of wheat in 2009 were China (115 million metric tonnes), India (81 MMT), Russian Federation (62 MMT), United States (60 MMT) and France (38 MMT). The wheat farm productivity in India and Russia were about 35% of the wheat farm productivity in France. China's farm productivity for wheat, in 2009, was about double that of Russia.
In addition to gaps in farming system technology and knowledge, some large wheat grain producing countries have significant losses after harvest at the farm and because of poor roads, inadequate storage technologies, inefficient supply chains and farmers' inability to bring the produce into retail markets dominated by small shopkeepers. Various studies in India, for example, have concluded that about 10% of total wheat production is lost at farm level, another 10% is lost because of poor storage and road networks, and additional amounts lost at the retail level. One study claims that if these post-harvest wheat grain losses could be eliminated with better infrastructure and retail network, in India alone enough food would be saved every year to feed 70 to 100 million people over a year.
Wheat futures are traded on the Chicago Board of Trade, Kansas City Board of Trade, and Minneapolis Grain Exchange, and have delivery dates in March (H), May (K), July (N), September (U), and December (Z).
|Source: UN Food & Agriculture Organization |
There are substantial differences in wheat farming, trading, policy, sector growth, and wheat uses in different regions of the world. In the EU and Canada for instance, there is significant addition of wheat to animal feeds, but less so in the United States.
The largest exporters of wheat in 2009 were, in order of exported quantities: United States, EU-27, Canada, Russian Federation, Australia, Ukraine and Kazakhstan. Upon the results of 2011, Ukraine became the world's sixth wheat exporter as well. The largest importers of wheat in 2009 were, in order of imported quantities: Egypt, EU-27, Brazil, Indonesia, Algeria and Japan. EU-27 was on both export and import list, because EU countries such as Italy and Spain imported wheat, while other EU-27 countries exported their harvest. The Black Sea region – which includes Kazakhstan, the Russian Federation and Ukraine – is amongst the most promising area for grain exporters; it possess significant production potential in terms of both wheat yield and area increases. The Black Sea region is also located close to the traditional grain importers in the Middle East, North Africa and Central Asia.
In the rapidly developing countries of Asia, westernization of diets associated with increasing prosperity is leading to growth in per capita demand for wheat at the expense of the other food staples.
In the past, there has been significant governmental intervention in wheat markets, such as price supports in the US and farm payments in the EU. In the EU these subsidies have encouraged heavy use of fertilizer inputs with resulting high crop yields. In Australia and Argentina direct government subsidies are much lower.
World's most productive wheat farms and farmers
The average annual world farm yield for wheat was 3.3 tonnes per hectare (330 grams per square meter), in 2013.
New Zealand wheat farms were the most productive in 2013, with a nationwide average of 9.1 tonnes per hectare. Ireland was a close second.
Various regions of the world hold wheat production yield contests every year. Yields above 12 tonnes per hectare are routinely achieved in many parts of the world. Chris Dennison of Oamaru, New Zealand, set a world record for wheat yield in 2003 at 15.015 tonnes per hectare (223 bushels/acre). In 2010, this record was surpassed by another New Zealand farmer, Michael Solari, with 15.636 tonnes per hectare (232.64 bushels/acre) at Otama, Gore.
Wheat normally needs between 110 and 130 days between sowing and harvest, depending upon climate, seed type, and soil conditions (winter wheat lies dormant during a winter freeze). Optimal crop management requires that the farmer have a detailed understanding of each stage of development in the growing plants. In particular, spring fertilizers, herbicides, fungicides, and growth regulators are typically applied only at specific stages of plant development. For example, it is currently recommended that the second application of nitrogen is best done when the ear (not visible at this stage) is about 1 cm in size (Z31 on Zadoks scale). Knowledge of stages is also important to identify periods of higher risk from the climate. For example, pollen formation from the mother cell, and the stages between anthesis and maturity are susceptible to high temperatures, and this adverse effect is made worse by water stress. Farmers also benefit from knowing when the 'flag leaf' (last leaf) appears, as this leaf represents about 75% of photosynthesis reactions during the grain filling period, and so should be preserved from disease or insect attacks to ensure a good yield.
Several systems exist to identify crop stages, with the Feekes and Zadoks scales being the most widely used. Each scale is a standard system which describes successive stages reached by the crop during the agricultural season.
There are many wheat diseases, mainly caused by fungi, bacteria, and viruses. Plant breeding to develop new disease-resistant varieties, and sound crop management practices are important for preventing disease. Fungicides, used to prevent the significant crop losses from fungal disease, can be a significant variable cost in wheat production. Estimates of the amount of wheat production lost owing to plant diseases vary between 10–25% in Missouri. A wide range of organisms infect wheat, of which the most important are viruses and fungi.
The main wheat-disease categories are:
- Seed-borne diseases: these include seed-borne scab, seed-borne Stagonospora (previously known as Septoria), common bunt (stinking smut), and loose smut. These are managed with fungicides.
- Leaf- and head- blight diseases: Powdery mildew, leaf rust, Septoria tritici leaf blotch, Stagonospora (Septoria) nodorum leaf and glume blotch, and Fusarium head scab.
- Crown and root rot diseases: Two of the more important of these are 'take-all' and Cephalosporium stripe. Both of these diseases are soil borne.
- Stem rust diseases: Caused by basidiomycete fungi e.g. Ug99
- Viral diseases: Wheat spindle streak mosaic (yellow mosaic) and barley yellow dwarf are the two most common viral diseases. Control can be achieved by using resistant varieties.
Wheat is used as a food plant by the larvae of some Lepidoptera (butterfly and moth) species including the flame, rustic shoulder-knot, setaceous Hebrew character and turnip moth. Early in the season, many species of birds, including the long-tailed widowbird, and rodents feed upon wheat crops. These animals can cause significant damage to a crop by digging up and eating newly planted seeds or young plants. They can also damage the crop late in the season by eating the grain from the mature spike. Recent post-harvest losses in cereals amount to billions of dollars per year in the United States alone, and damage to wheat by various borers, beetles and weevils is no exception. Rodents can also cause major losses during storage, and in major grain growing regions, field mice numbers can sometimes build up explosively to plague proportions because of the ready availability of food. To reduce the amount of wheat lost to post-harvest pests, Agricultural Research Service scientists have developed an "insect-o-graph," which can detect insects in wheat that are not visible to the naked eye. The device uses electrical signals to detect the insects as the wheat is being milled. The new technology is so precise that it can detect 5-10 infested seeds out of 300,000 good ones. Tracking insect infestations in stored grain is critical for food safety as well as for the marketing value of the crop.
- Deficit irrigation
- Wheat germ oil
- Wheat production in the United States
- Wheat middlings
- Whole wheat flour
- Belderok, Robert 'Bob'; Mesdag, Hans; Donner, Dingena A (2000), Bread-Making Quality of Wheat, Springer, p. 3, ISBN 0-7923-6383-3
- Shewry, Peter R (2009), "Wheat", Journal of Experimental Botany 60 (6): 1537–1553, doi:10.1093/jxb/erp058
- James D. Mauseth (2014). Botany. Jones & Bartlett Publishers. p. 223. ISBN 978-1-4496-4884-8.
Perhaps the simplest of fruits are those of grasses (all cereals such as corn and wheat)...These fruits are caryopses.
- "FAOStat". Retrieved 27 January 2015.
- "World Wheat, Corn and Rice". Oklahoma State University, FAO Stat. Archived from the original on 10 June 2015.
- Curtis; Rajaraman; MacPherson (2002). "Bread Wheat". Food and Agriculture Organization of the United Nations.
- "Nutrient data laboratory". United States Department of Agriculture.
- Tanno, K Willcox; Willcox, G (2006). "How fast was wild wheat domesticated?". Science 311 (5769): 1886. doi:10.1126/science.1124635. PMID 16574859.
- Feldman, Moshe and Kislev, Mordechai E., Israel Journal of Plant Sciences, Volume 55, Number 3 - 4 / 2007, pp. 207 - 221, Domestication of emmer wheat and evolution of free-threshing tetraploid wheat in "A Century of Wheat Research-From Wild Emmer Discovery to Genome Analysis", Published Online: 3 November 2008
- Colledge, Sue; University College, London. Institute of Archaeology (2007). The origins and spread of domestic plants in southwest Asia and Europe. Left Coast Press. pp. 40–. ISBN 978-1-59874-988-5. Retrieved 5 July 2011.
- C. Michael Hogan. 2013. Wheat. Encyclopedia of Earth. National Council of Science and the Environment. ed. Lakhdar Boukerrou
- Heun, MR; et al. (1997). "Site of Einkorn Wheat Domestication Identified by DNA Fingerprinting". Science 278: 1312–4. doi:10.1126/science.278.5341.1312.
- Ozkan, H; Brandolini, A; Schäfer-Pregl, R; Salamini, F (October 2002). "AFLP analysis of a collection of tetraploid wheats indicates the origin of emmer and hard wheat domestication in southeast Turkey". Molecular Biology and Evolution 19 (10): 1797–801. doi:10.1093/oxfordjournals.molbev.a004002. PMID 12270906.
- Diamond J (1997) Guns, Germs and Steel, A short history of everybody for the last 13,000 years. Viking UK Random House ISBN 0-09-930278-0
- Direct quotation: Grundas ST: Chapter: Wheat: The Crop, in Encyclopedia of Food Sciences and Nutrition p6130, 2003; Elsevier Science Ltd
- "the science in detail – Wheats DNA – Research – Archaeology – The University of Sheffield". Sheffield.ac.uk. 19 July 2011. Retrieved 27 May 2012.
- Belderok B et al. (2000) Bread-Making Quality of Wheat Springer p 3 ISBN 0-7923-6383-3
- Abengoa And Dyadic Sign Ethanol R&D Agreement Posted 31 October 2006
- Cauvain SP, Cauvain P (2003) Bread Making CRC Press p 540 ISBN 1-85573-553-9
- Bergen R 'American wheat beers' In Brewing Techniques
- FAOSTAT Agricultural statistics 2005 data values
- Hancock, James F. (2004) Plant Evolution and the Origin of Crop Species. CABI Publishing. ISBN 0-85199-685-X.
- Hoisington, D; Khairallah, M; Reeves, T; Ribaut, JM; Skovmand, B; Taba, S; Warburton, M (1999). "Plant genetic resources: What can they contribute toward increased crop productivity?". Proc Natl Acad Sci USA 96 (11): 5937–43. doi:10.1073/pnas.96.11.5937. PMC 34209. PMID 10339521.
- Basra, AS (1999) Heterosis and Hybrid Seed Production in Agronomic Crops Haworth Press pp 81-82 ISBN 1-56022-876-8
- (12 May 2013) Cambridge-based scientists develop 'superwheat' BBC News UK, Retrieved 25 May 2013
- Synthetic hexaploids
- (2013) Synthetic hexaploid wheat UK National Institute of Agricultural Botany, Retrieved 25 May 2013
- Drysdale wheat bred for dry conditions
- Huge potential for water-efficient wheat
- Condon, AG; Farquhar, GD; Richards, RA (1990). "Genotypic variation in carbon isotope discrimination and transpiration efficiency in wheat. Leaf gas exchange and whole plant studies". Australian Journal of Plant Physiology 17: 9–22. doi:10.1071/PP9900009.
- BBSRC press release UK researchers release draft sequence coverage of wheat genome BBSRC, 27 August 2010
- UK scientists publish draft sequence coverage of wheat genome
- Hall. "Analysis of the bread wheat genome using whole-genome shotgun sequencing: Nature : Nature Publishing Group". Nature. Retrieved 5 February 2014.
- Bajaj, Y. P. S. (1990) Wheat. Springer. pp. 161-63. ISBN 3-540-51809-6.
- Joint FAO/IAEA META Information Portal
- Verma, Shailender Kumar; Kumar, Satish; Sheikh, Imran; Malik, Sachin; Mathpal, Priyanka; Chugh, Vishal; Kumar, Sundip; Prasad, Ramasare; Dhaliwal, Harcharan Singh (2016-03-03). "Transfer of useful variability of high grain iron and zinc from Aegilops kotschyi into wheat through seed irradiation approach". International Journal of Radiation Biology 92 (3): 132–139. doi:10.3109/09553002.2016.1135263. ISSN 0955-3002. PMID 26883304.
- Mike Abram for Farmers' Weekly. May 17, 2011. Hybrid wheat to make a return
- Bill Spiegel for agriculture.com March 11, 2013 Hybrid wheat's comeback
- History of hybrid wheat
- Basra, Amarjit S. (1999) Heterosis and Hybrid Seed Production in Agronomic Crops. Haworth Press. pp. 81-82. ISBN 1-56022-876-8.
- Potts, D. T. (1996) Mesopotamia Civilization: The Material Foundations Cornell University Press. p. 62. ISBN 0-8014-3339-8.
- Nevo, Eviatar & A. B. Korol & A. Beiles & T. Fahima. (2002) Evolution of Wild Emmer and Wheat Improvement: Population Genetics, Genetic Resources, and Genome.... Springer. p. 8. ISBN 3-540-41750-8.
- Vaughan, J. G. & P. A. Judd. (2003) The Oxford Book of Health Foods. Oxford University Press. p. 35. ISBN 0-19-850459-4.
- Bridgwater, W. & Beatrice Aldrich. (1966) The Columbia-Viking Desk Encyclopedia. Columbia University. p. 1959.
- Moon, David (2008). "In the Russian Steppes: the Introduction of Russian Wheat on the Great Plains of the UNited States". Journal of Global History 3: 203–225. doi:10.1017/s1740022808002611.
- "National Nutrient Database for Standard Reference Release 28". United States Department of Agriculture: Agricultural Research Service.
- "Nutrition facts, calories in food, labels, nutritional information and analysis". NutritionData.com.
- "USDA Table of Nutrient Retention Factors, Release 6" (PDF). USDA. USDA. Dec 2007.
- "Nutritional Effects of Food Processing". NutritionData.com.
- USDA National Nutrient Database for Standard Reference, Release 25 (2012)
- "FAOStat". Retrieved 27 January 2015.
- "USA: U.S., Australia, India partnership to develop climate-resilient varieties of rice and wheat :: Agriculture in the Black Sea Region". Bs-agro.com. 24 May 2013. Retrieved 5 February 2014.
- Preedy, Victor; et al. (2011). Nuts and seeds in health and disease prevention. Academic Press. pp. 960–967. ISBN 978-0-12-375688-6.
- Qin Liu; et al. (2010). "Comparison of Antioxidant Activities of Different Colored Wheat Grains and Analysis of Phenolic Compounds". Journal of Agricultural and Food Chemistry 58 (16): 9235–9241. doi:10.1021/jf101700s.
- Lundin KE, Wijmenga C (Sep 2015). "Coeliac disease and autoimmune disease-genetic overlap and screening". Nat Rev Gastroenterol Hepatol 12 (9): 507–15. doi:10.1038/nrgastro.2015.136. PMID 26303674.
- Fasano A (Apr 2005). "Clinical presentation of celiac disease in the pediatric population". Gastroenterology 128 (4 Suppl 1): S68–73. doi:10.1053/j.gastro.2005.02.015. PMID 15825129.
- Elli L, Branchi F, Tomba C, Villalta D, Norsa L, Ferretti F, Roncoroni L, Bardella MT (Jun 2015). "Diagnosis of gluten related disorders: Celiac disease, wheat allergy and non-celiac gluten sensitivity". World J Gastroenterol 21 (23): 7110–9. doi:10.3748/wjg.v21.i23.7110. PMC 4476872. PMID 26109797.
- Catassi C, Bai J, Bonaz B, Bouma G, Calabrò A, Carroccio A, Castillejo G, Ciacci C, Cristofori F, Dolinsek J, Francavilla R, Elli L, Green P, Holtmeier W, Koehler P, Koletzko S, Meinhold C, Sanders D, Schumann M, Schuppan D, Ullrich R, Vécsei A, Volta U, Zevallos V, Sapone A, Fasano A (2013). "Non-celiac gluten sensitivity: the new frontier of gluten related disorders". Nutrients (Review) 5 (10): 3839–3853. doi:10.3390/nu5103839. ISSN 2072-6643. PMID 24077239.
- Ludvigsson JF, Leffler DA, Bai JC, Biagi F, Fasano A, Green PH, Hadjivassiliou M, Kaukinen K, Kelly CP, Leonard JN, Lundin KE, Murray JA, Sanders DS, Walker MM, Zingone F, Ciacci C (January 2013). "The Oslo definitions for coeliac disease and related terms". Gut 62 (1): 43–52. doi:10.1136/gutjnl-2011-301346. PMC 3440559. PMID 22345659.
- Molina-Infante J, Santolaria S, Sanders DS, Fernández-Bañares F (May 2015). "Systematic review: noncoeliac gluten sensitivity". Aliment Pharmacol Ther 41 (9): 807–20. doi:10.1111/apt.13155. PMID 25753138.
- "USDA National Nutrient Database for Standard Reference". United States Department of Agriculture. Archived from the original on 3 March 2015.
- "Nutrient data laboratory". United States Department of Agriculture. Retrieved June 2014.
- Ali, MB (2002), Characteristics and production costs of US wheat farms, USDA, SB-974-5 ERS
- Long, Victoria Sizemore (28 September 2007), "Wheat futures again hit new highs", The Kansas City Star (article), archived from the original on 20 October 2007
- CIMMYT World wheat facts and trends 1998-9.
- Kiss, Istvan. "Significance of wheat production in world economy and position of Hungary in it" (PDF). Agroinform Publishing House, Budapest, Hungary. Retrieved 2 February 2013.
- See Chapter 1, Slafer GA, Satorre EH (1999) Wheat: Ecology and Physiology of Yield Determination Haworth Press Technology & Industrial ISBN 1-56022-874-1.
- Swaminathan MS (2004) Stocktake on cropping and crop science for a diverse planet
- Umbers, Alan (2006, Grains Council of Australia Limited) Grains Industry trends in Production - Results from Today's Farming Practices
- Basavaraja, H.; et al. "Economic Analysis of Post-harvest Losses in Food Grains in India: A Case Study of Karnataka" (PDF). Agricultural Economics Research Review 20: 117–126.
- List of Commodity Delivery Dates on Wikinvest
- "Production of Wheat by countries". UN Food & Agriculture Organization (FAO). 2011. Retrieved 26 January 2015.
- "Wheat Flour: Agri Handbook" (PDF). Food and Agriculture Organization of the United Nations. 2011. pp. 12–18.
- Ukraine becomes world's third biggest grain exporter in 2011 — Minister, Black sea grain, archived from the original on 10 October 2014
- "World Wheat Overview and Outlook 2000–01", Facts & trends (research), CIMMYT, archived from the original on 22 April 2009
- "FAOSTAT: Production-Crops, 2013 data". Food and Agriculture Organization of the United Nations. 2013.
- Barker, Bruce (2011). "Breaking the Guinness world record for wheat yield". Top Crop Manager.
- Slafer GA, Satorre EH (1999) Wheat: Ecology and Physiology of Yield Determination Haworth Press Technology & Industrial ISBN 1-56022-874-1. pp 322-3
- Crop Disease Management Bulletin 631-98. Wheat Diseases[dead link]
- "G4319 Wheat Diseases in Missouri, MU Extension". Muextension.missouri.edu. Archived from the original on 27 February 2007. Retrieved 18 May 2009.
- C.Michael Hogan. 2013. Wheat. Encyclopedia of Earth, National Council for Science and the Environment, Washington DC ed. P. Saundry
- Gautam, P. and R. Dill-Macky. 2012. Impact of moisture, host genetics and Fusarium graminearum isolates on Fusarium head blight development and trichothecene accumulation in spring wheat. Mycotoxin Research Vol 28 Iss 1 doi:10.1007/s12550-011-0115-6 
- Biological Control of Stored-Product Pests. Biological Control News Volume II, Number 10 October 1995
- CSIRO Rodent Management Research Focus: Mice plagues
- "ARS, Industry Cooperation Yields Device to Detect Insects in Stored Wheat". USDA Agricultural Research Service. 24 June 2010.
This article incorporates material from the Citizendium article "Wheat", which is licensed under the Creative Commons Attribution-ShareAlike 3.0 Unported License but not under the GFDL.
- Bonjean, A.P., and W.J. Angus (editors). The World Wheat Book: a history of wheat breeding. Lavoisier Publ., Paris. 1131 pp. (2001). ISBN 2-7430-0402-9
- Christen, Olaf, ed. (2009) (in German), Winterweizen. Das Handbuch für Profis, DLG-Verlags-GmbH, ISBN 978-3-7690-0719-0
- Garnsey Peter, Grain for Rome, in Garnsey P., Hopkins K., Whittaker C. R. (editors), Trade in the Ancient Economy, Chatto & Windus, London 1983
- Head L., Atchison J., and Gates A. Ingrained: A Human Bio-geography of Wheat. Ashgate Publ., Burlington. 246 pp. (2012). ISBN 978-1-4094-3787-1
- Jasny Naum, The daily bread of ancient Greeks and Romans, Ex Officina Templi, Brugis 1950
- Jasny Naum, The Wheats of Classical Antiquity, J. Hopkins Press, Baltimore 1944
- Heiser Charles B., Seed to civilisation. The story of food, (Harvard University Press, 1990)
- Harlan Jack R., Crops and man, American Society of Agronomy, Madison 1975
- Padulosi, S.; Hammer, K.; Heller, J., eds. (1996). Hulled wheats. Promoting the conservation and use of underutilized and neglected crops. 4. International Plant Genetic Resources Institute, Rome, Italy.[dead link]
- Saltini Antonio, I semi della civiltà. Grano, riso e mais nella storia delle società umane, Prefazione di Luigi Bernabò Brea, Avenue Media, Bologna 1996
- Sauer Jonathan D., Geography of Crop Plants. A Select Roster, CRC Press, Boca Raton
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