Temporal range: Late Pliocene to Recent
|A South African cheetah (A. jubatus jubatus)|
|The range of the cheetah|
The cheetah (Acinonyx jubatus) is a large feline (family Felidae, subfamily Felinae) inhabiting most of Africa and parts of Iran. It is the only extant member of the genus Acinonyx. The cheetah can run faster than any other land animal— as fast as 110 to 120 km/h (68 to 75 mph) in short bursts covering distances up to 500 m (1,600 ft), and has the ability to accelerate from 0 to 96 km/h (60 mph) in three seconds.
The cheetah is a unique felid, with its closest living relatives being the puma and jaguarundi of the Americas. This cat is notable for modifications in the species' paws, being one of the few felids with only semi-retractable claws.
Its main hunting strategy is to trip swift prey such as various antelope species and hares with their dewclaw. Almost every facet of the cheetah's anatomy has evolved to maximise its success in the chase, the result of an evolutionary arms race with its prey. Due to this specialisation, however, the cheetah is poorly equipped to defend itself against other large predators, with speed being its main means of defence.
In the wild, the cheetah is a prolific breeder, with up to nine cubs in a litter. The majority of cubs do not survive to adulthood, mainly as a result of depredation from other predators. The rate of cub mortality varies from area to area, from 50% to 75%, and in extreme cases such as the Serengeti ecosystem, up to 90%. Cheetahs are notoriously poor breeders in captivity, though several organizations, such as the De Wildt Cheetah and Wildlife Centre, have succeeded in breeding high numbers of cubs.
The cheetah is listed as vulnerable, facing various threats including loss of habitat and prey, conflict with humans, the illegal pet trade, competition with and predation by other carnivores, and a gene pool with very low variability. It is a charismatic species and many captive cats are "ambassadors" for their species and wildlife conservation in general.
- 1 Etymology
- 2 Genetics, evolution, and classification
- 3 Description
- 4 Range and habitat
- 5 Reproduction and behavior
- 6 Relationship with humans
- 7 In popular culture
- 8 References
- 9 Sources
- 10 Further reading
- 11 External links
Genetics, evolution, and classification
The cheetah has unusually low genetic variability. This is accompanied by a very low sperm count, motility, and deformed flagella. Skin grafts between unrelated cheetahs illustrate the former point, in that there is no rejection of the donor skin. It is thought that the species went through a prolonged period of inbreeding following a genetic bottleneck during the last ice age. This suggests that genetic monomorphism did not prevent the cheetah from flourishing across two continents for thousands of years.
The cheetah likely evolved in Africa during the Miocene epoch (26 million to 7.5 million years ago), before migrating to Asia. Recent research has placed the last common ancestor of all existing populations as living in Asia 11 million years ago, which may lead to revision and refinement of existing ideas about cheetah evolution.
The now-extinct species include Acinonyx pardinensis (Pliocene epoch), much larger than the modern cheetah and found in Europe, India, and China; and Acinonyx intermedius (mid-Pleistocene period), found over the same range. The extinct genus Miracinonyx was extremely cheetah-like, but recent DNA analysis has shown that Miracinonyx inexpectatus, Miracinonyx studeri, and Miracinonyx trumani (early to late Pleistocene epoch), found in North America and called the "North American cheetah" are not true cheetahs, instead being close relatives to the cougar.
Although many sources list six or more subspecies of cheetah, the taxonomic status of most of these subspecies is unresolved. Acinonyx rex—the king cheetah—was abandoned as a species after it was discovered that the variation was caused by a single recessive gene. The subspecies Acinonyx jubatus guttatus, the woolly cheetah, may also have been a variation due to a recessive gene. Some of the most commonly recognized subspecies include:
|South African cheetah (A. j. jubatus), also called the Namibian cheetah||Lives in South Africa, Namibia, Zimbabwe, and Botswana, and is the most common subspecies. In 2007, there were 1,800 in Botswana, 550-850 in South Africa, 400 in Zimbabwe, 100 in Zambia, more than 50-90 in Mozambique and more than 25-50 in Malawi. In Namibia, the population has increased from 2,500 to 3,500 today. It lives in grasslands, savannahs, arid environments, open fields and mountains, and occupies a medium size range among surviving subspecies.|
|Tanzanian cheetah (A. j. raineyii), also commonly known as East African cheetah||Is found in Kenya, Somalia, Tanzania, and Uganda. The total population in 2007 was estimated at 2,572 adults and independent adolescents. Tanzanian cheetahs are the second-common subspecies after the most numerous South African cheetah. It is the largest subspecies.|
|Sudan cheetah (A. j. soemmeringii), also known as Central or Northeast African cheetah||Found in the central and northeastern regions of the continent and in the Horn of Africa, this subspecies was considered identical to the South African cheetah until a 2011 genetic analysis demonstrated significant differences. It is the second-largest of the surviving subspecies. In 2002, the total population was estimated at around 2,000 individuals in the wild.
Northeastern Africa: (Djibouti, Egypt, Eritrea, Ethiopia, Libya, Somalia, South Sudan and Sudan)
|Northwest African cheetah (A. j. hecki), also known as the Saharan cheetah||Lives in the northwestern part of Africa. With an estimated total world population of only 250 mature individuals, it is listed as critically endangered. It is the palest and smallest African cheetah subspecies.|
|Asiatic cheetah (A. j. venaticus), also known as Iranian or Indian cheetah||Found only on the deserts of Iran, and is thus the only surviving cheetah subspecies indigenous to Asia. It is the most critically endangered subspecies of cheetah, and one of the most endangered animals in the world. As of 2015, the wild population is estimated at approximately 50 to 70, found mostly in Iran's national parks. It is among the smallest of the cheetahs, with a slighter build than the African cheetahs, more fur on the back of the neck, a longer and more powerful neck, thinner tear marks and a smaller head. It is the only subspecies to possess a woolly winter coat.
Western Asia: (Georgia, Iran, Iraq, Israel, Jordan, Kuwait, Lebanon, Oman, Saudi Arabia, Syria, Turkey, United Arab Emirates and Yemen)
The cheetah's chest is deep and its waist is narrow. The coarse, short fur of the cheetah is tan with round black spots measuring from 2 to 3 cm (0.79 to 1.18 in) across, affording it some camouflage while hunting. There are no spots on its white underside, but the tail has spots, which merge to form four to six dark rings at the end. The tail usually ends in a bushy white tuft. The cheetah has a small head with high-set eyes. Black "tear marks" running from the corner of its eyes down the sides of the nose to its mouth keep sunlight out of its eyes and aid in hunting and seeing long distances. Its thin and fragile body make it well-suited to short bursts of high speed, but not to long-distance running.
Agility, rather than raw speed, accounts for much of the cheetah's ability to catch prey. Cheetahs can accelerate four times as fast as a human (thanks to greater muscle power) and can slow down by 14 kilometers per hour in one stride. They can hunt successfully in densely vegetated areas.
The adult cheetah weighs from 21 to 72 kg (46 to 159 lb). Its total head-and-body length is from 110 to 140 cm (43 to 55 in), while the tail can measure 60 to 84 cm (24 to 33 in) in length. Cheetahs are 66 to 94 cm (26 to 37 in) tall at the shoulder. Males tend to be slightly larger than females and have slightly bigger heads, but there is not a great variation in cheetah sizes and it is difficult to tell males and females apart by appearance alone. Compared to a similarly sized leopard, the cheetah is generally shorter-bodied, but is longer tailed and taller (it averages about 90 cm (35 in) tall) and so it appears more streamlined.
Some cheetahs have a rare fur pattern mutation of larger, blotchy, merged spots. Known as "king cheetahs," they were once thought to constitute a separate subspecies but are in fact African cheetahs; their unusual fur pattern is the result of a single recessive gene. The "king cheetah" has only been seen in the wild a handful of times, but it has been bred in captivity.
The cheetah's paws have semi-retractable claws (known only in three other cat species: the fishing cat, the flat-headed cat and the Iriomote cat), offering extra grip in its high-speed pursuits. The ligament structure of the cheetah's claws is the same as those of other cats; it simply lacks the sheath of skin and fur present in other varieties, and therefore, with the exception of the dewclaw, the claws are always visible. The dewclaw is much shorter and straighter than that of other cats.
Adaptations that enable the cheetah to run as fast as it does include large nostrils that allow for increased oxygen intake, and an enlarged heart and lungs that work together to circulate oxygen efficiently. During a typical chase, its respiratory rate increases from 60 to 150 breaths per minute. While running, in addition to having good traction due to its semi-retractable claws, the cheetah uses its tail as a rudder-like means of steering to allow it to make sharp turns, necessary to outflank prey animals that often make such turns to escape.
Unlike true big cats of subfamily Pantherinae, the cheetah can purr as it inhales, but cannot roar. By contrast, the big cats can roar but cannot purr, except while exhaling. The cheetah is still considered by some to be the smallest of the big cats. While it is often mistaken for the leopard, the cheetah does have distinguishing features, such as the aforementioned long "tear-streak" lines that run from the corners of its eyes to its mouth, and spots that are not "rosettes". The thinner body frame of the cheetah is also very different from that of the leopard.
The cheetah is a vulnerable species. Of all the big cats, it is the least able to adapt to new environments. It has always proved difficult to breed in captivity, although recently a few zoos have managed to succeed at this. One technique has been to introduce a dog as a playmate and guard dog to enable a captive cheetah to feel less threatened.
Once widely hunted for its fur, the cheetah now suffers more from the loss of both habitat and prey, conflict with humans and the illegal pet trade.
The cheetah was formerly considered to be particularly primitive among the cats and to have evolved approximately 18 million years ago. However, new research suggests the last common ancestor of all 40 existing species of felines lived more recently than about 11 million years ago. The same research indicates that the cheetah, while highly derived morphologically, is not of particularly ancient lineage, having separated from its closest living relatives (Puma concolor, the cougar, and Puma yaguarondi, the jaguarundi) around five million years ago. These felids have not changed appreciably since they first appeared in the fossil record.
Morphs and variations
The king cheetah is a rare mutation of the cheetah characterized by a distinct fur pattern. It was first noted in what was then Southern Rhodesia (modern-day Zimbabwe) in 1926. In 1927, the naturalist Reginald Innes Pocock declared it a separate species, but reversed this decision in 1939 due to lack of evidence; but in 1928, a skin purchased by Walter Rothschild was found to be intermediate in pattern between the king cheetah and spotted cheetah and Abel Chapman considered it to be a color form of the spotted cheetah. Twenty-two such skins were found between 1926 and 1974. Since 1927, the king cheetah was reported five more times in the wild. Although strangely marked skins had come from Africa, a live king cheetah was not photographed until 1974 in South Africa's Kruger National Park. Cryptozoologists Paul and Lena Bottriell photographed one during an expedition in 1975. They also managed to obtain stuffed specimens. It appeared larger than a spotted cheetah and its fur had a different texture. There was another wild sighting in 1986—the first in seven years. By 1987, thirty-eight specimens had been recorded, many from pelts.
Its species status was resolved in 1981 when king cheetahs were born at the De Wildt Cheetah and Wildlife Centre in South Africa. In May 1981, two spotted sisters gave birth there and each litter contained one king cheetah. The sisters had both mated with a wild-caught male from the Transvaal area (where king cheetahs had been recorded). Further king cheetahs were later born at the Centre. It has been known to exist in Zimbabwe, Botswana and in the northern part of South Africa's Transvaal province.
In 2012, the cause of this alternative coat pattern was found to be a mutation in the gene for transmembrane aminopeptidase Q (Taqpep), the same gene responsible for the striped "mackerel" versus blotchy "classic" patterning seen in tabby cats. The mutation is recessive and must be inherited from both parents for this pattern to appear, which is one reason why it is so rare.
Other color variations
Other rare color morphs of the species include speckles, melanism, albinism, abundism, chocolate, erythrism, strawberry, isabelline, golden, Maltese, chinchilla, black-marked, red-cream marked, ticked, charcoal, mosaicism, leucism, lavender-marked, piebaldism, and flavism. Most have been reported in Indian cheetahs, particularly in captive specimens kept for hunting.
The Mughal Emperor of India, Jahangir, recorded having a white cheetah presented to him in 1608. In the memoirs of Tuzk-e-Jahangiri, the Emperor, says that in the third year of his reign, "Raja Bir Singh Deo brought a white cheetah to show me. Although other sorts of creatures, both birds and beasts have white varieties ... I had never seen a white cheetah. Its spots, which are (usually) black, were of a blue color, and the whiteness of the body also inclined to bluishness." This suggests a chinchilla mutation which restricts the amount of pigment on the hair shaft. Although the spots were formed of black pigment, the less dense pigmentation gives a hazy, grayish effect. As well as Jahangir's white cheetah at Agra, a report of "incipient albinism" has come from Beaufort West according to Guggisberg.
In a letter to "Nature in East Africa", H. F. Stoneham reported a melanistic cheetah (black with ghost markings) in the Trans-Nzoia District of Kenya in 1925. Vesey Fitzgerald saw a melanistic cheetah in Zambia in the company of a spotted cheetah. Red (erythristic) cheetahs have dark tawny spots on a golden background. Cream (isabelline) cheetahs have pale red spots on a pale background. Some desert region cheetahs are unusually pale; probably they are better-camouflaged and therefore better hunters and more likely to breed and pass on their paler colouration. Blue (Maltese or grey) cheetahs have variously been described as white cheetahs with grey-blue spots (chinchilla) or pale grey cheetahs with darker grey spots (Maltese mutation). A ticked was shot in Tanzania in 1921 (Pocock); it had only a few spots on the neck and back, and these were unusually small. Another ticked cheetah color-morph was photographed in Kenya in 2012.
Range and habitat
There are several geographically isolated populations of cheetah, all of which are found in 23 African countries. A small population (estimated at about 50) survive in the Khorasan Province of Iran, where conservationists are taking steps to protect them.
It is possible, though doubtful, that some cheetahs remain in India. There have also been several unconfirmed reports of Asiatic Cheetahs in the Balochistan province of Pakistan, with at least one dead animal being discovered recently.
The cheetah thrives in areas with vast expanses of land where prey is abundant. The cheetah likes to live in an open biotope, such as semidesert, prairie, and thick brush, though it can be found in a variety of habitats. In Namibia, for example, it lives in grasslands, savannahs, areas of dense vegetation, and mountainous terrain.
Reproduction and behavior
Females reach maturity in twenty to twenty-four months, and males around twelve months (although they do not usually mate until at least three years old), and mating occurs throughout the year. A study of cheetahs in the Serengeti showed females are sexually promiscuous and often have cubs by many different males.
Females give birth to up to nine cubs after a gestation period of ninety to ninety-eight days, although the average litter size is four. Cubs weigh from 150 to 300 g (5.3 to 10.6 oz) at birth. Unlike some other cats, the cheetah is born with its characteristic spots. Cubs are also born with a downy underlying fur on their necks, called a mantle, extending to mid-back. This gives them a mane or Mohawk-type appearance; this fur is shed as the cheetah grows older. It has been speculated that this mane gives a cheetah cub the appearance of the honey badger (ratel), to scare away potential aggressors. The mantle is also believed to act as camouflage to hide the cubs from predators in tall grass. Life span is up to twelve years in the wild, but up to twenty years in captivity. The rate of cub mortality varies from area to area, from 50% to 75%, and in extreme cases such as the Serengeti ecosystem, up to 90%. In comparison to the Serengeti, the survival rate of cheetah cubs in the Kgalagadi area was seven times higher.
Unlike males, females are solitary and tend to avoid each other, though some mother/daughter pairs have been known to be formed for small periods of time. The cheetah has a unique, well-structured social order. Females live alone, except when they are raising cubs and they raise their cubs on their own. The first eighteen months of a cub's life are important; cubs must learn many lessons, because survival depends on knowing how to hunt wild prey species and avoid other predators. At 18 months, the mother leaves the cubs, who then form a sibling ("sib") group that will stay together for another six months. At about two years, the female siblings leave the group, and the young males remain together for life.
Males are often social and may group together for life, usually with their brothers in the same litter; although if a cub is the only male in the litter then two or three lone males may form a group, or a lone male may join an existing group. These groups are called coalitions. In the Serengeti, it was found that 41% of the adult males were solitary, 40% lived in pairs and 19% lived in trios.
A coalition is six times more likely to obtain an animal territory than a lone male, although studies have shown that coalitions keep their territories just as long as lone males— between four to four and a half years.
Males are territorial. Females' home ranges can be very large and a territory including several females' ranges is impossible to defend. Instead, males choose the points at which several of the females' home ranges overlap, creating a much smaller space, which can be properly defended against intruders while maximizing the chance of reproduction. Coalitions will try their best to maintain territories to find females with whom they will mate. The size of the territory also depends on the available resources; depending on the habitat, the size of a male's territory can vary greatly from 37 to 160 km2 (14 to 62 sq mi).
Scent is an important means of communication among cheetahs. Males mark their territory by urinating or defecating on objects that stand out, such as trees, logs, or termite mounds. When male cheetahs urine-mark their territories, they stand less than one meter away from a tree or rock surface with the tail raised, pointing the penis either horizontally backward or 60° upward. Male coalitions are able to defend the best territories through joint scent-marking. Males will attempt to kill any intruders, and fights result in serious injury or death.
Unlike males and other felines, females do not establish territories. Instead, the area they live in is termed a home range. These overlap with other females' home ranges, often those of their daughters, mothers, or sisters. Females always hunt alone, although cubs will accompany their mothers to learn to hunt once they reach the age of five to six weeks.
The size of a home range depends entirely on the availability of prey. Cheetahs in southern African woodlands have ranges as small as 34 km2 (13 sq mi), while in some parts of Namibia they can reach 1,500 km2 (580 sq mi).
The cheetah cannot roar, but ranks among the more vocal felids. Several sources refer to a wide variety of cheetah vocalizations, but most of these lack a detailed acoustic description which makes it difficult to reliably assess exactly what terms refer to exactly what vocalizations. A short review of the terminology encountered is found in. Some of the vocalizations listed in the literature are:
- Chirping: When a cheetah attempts to find another, or a mother tries to locate her cubs, it uses a high-pitched barking called chirping. The chirps made by a cheetah cub sound more like a bird chirping, and so are termed chirping, too.
- Churring or stuttering: This vocalization is emitted by a cheetah during social meetings. A churr can be seen as a social invitation to other cheetahs, an expression of interest, uncertainty, or appeasement or during meetings with the opposite sex (although each sex churrs for different reasons).
- Growling: This vocalization is often accompanied by hissing and spitting and is exhibited by the cheetah during annoyance, or when faced with danger.
- Yowling: This is an escalated version of growling, usually displayed when danger worsens.
- Agonistic vocalizations: a combination of growls, moans, hisses and the "trademark" cheetah spit, which is most often accompanied by a forceful "paw hit" on the ground.
- Purring: This is made when the cheetah is content, usually during pleasant social meetings (mostly between cubs and their mothers). A characteristic of purring is that it is realized on both egressive and ingressive airstream, as seen and heard on online video and audio.
Diet and hunting
The cheetah is a carnivore, eating mostly mammalian herbivores under 40 kg (88 lb) and that which specialise in eating C3 plants, including the Thomson's gazelle, the Grant's gazelle, the springbok, impala and blesbok. The young of larger mammals such as wildebeests and zebras are taken at times, and adults too, when cheetahs hunt in groups. Guineafowl and hares are also prey. Ostriches are also taken on occasion. In Iran, cheetahs prey on the Chinkara, Goitered gazelle, ibexes and wild sheep.
While the other big cats often hunt by night, the cheetah is a diurnal hunter. It will, however, hunt on moonlit nights during the full Moon as well, where visibility is excellent. It hunts usually either early in the morning or later in the evening when it is not too hot, but there is still enough light.
The cheetah hunts by vision rather than by scent. Prey is stalked to within 10–30 m (33–98 ft), then chased. This is usually over in less than a minute, and if the cheetah fails to make a catch quickly, it will give up. The cheetah has an average hunting success rate of around 50%. Cheetahs can run at a very high speed; in just two seconds they can accelerate from 0 to 75 kilometers per hour. The estimated top speed of the cheetah ranges from 90 to 128 kilometers per hour. Cheetahs refuse to run when their body temperature reaches 40.5 °C.
Running at such high speeds puts a great deal of strain on the cheetah's body. When sprinting, the cheetah's respiratory rate goes from 60 to 150 breaths per minute, while their body temperature has been measured at 40.5 °C (over 2 °C above normal) during a 375-meter sprint. If it is a hard chase, it sometimes needs to rest for half an hour or more.
The cheetah kills its prey by tripping it during the chase, then biting it on the underside of the throat to suffocate it; the cheetah is not strong enough to break the necks of most prey. The bite may also puncture a vital artery in the neck. Then the cheetah proceeds to devour its catch as quickly as possible before the kill is taken by stronger predators.
Data from 367 runs by three female and two male adults, with an average run distance of 173 m, showed that hunting cheetahs can run 58 miles (93 km) per hour. A recent study that followed five African cheetahs indicated that cheetahs relied most heavily on acceleration. Most chases involved extreme maneuverability more than speed. The study indicated that cheetahs seemed to rarely run close to 60 mph or more; on most hunts they reached 30 to 35 mph, but they accelerated and changed direction much more rapidly than any other land animal.
The diet of a cheetah depends on the area in which it lives. For example, on the East African plains, its preferred prey is the Thomson's gazelle. This small antelope is smaller than the cheetah, which makes it an appropriate prey. In contrast, in Kwa-Zulu Natal, the main species of the cheetah's prey preference is the significantly larger nyala, which can weigh up to 130 kg (290 lb) in the male. Cheetahs look for individuals that have strayed some distance from their group, and do not necessarily seek out old or weak ones.
Interspecific predatory relationships
Despite their speed and hunting prowess, cheetahs are largely outranked by other large predators in most of their range. Because they have evolved for short bursts of extreme speed at the expense of their power, they cannot defend themselves against most of Africa's other predator species. They usually avoid fighting and will surrender a kill immediately to even a single hyena, rather than risk injury. Because cheetahs rely on their speed to obtain their meals, any injury that slows them down could essentially be life-threatening.
A cheetah has a 50% chance of losing its kill to other predators. Cheetahs avoid competition by hunting at different times of the day and by eating immediately after the kill. Due to the reduction in habitat in Africa, cheetahs in protected areas face greater pressure from other larger predators, causing them to live outside of reserves and increasingly coming into conflict with humans. In Namibia, where the largest population of wild cheetah lives, 90% of these cheetahs live on farmland.
The cheetah's mortality is very high during the early weeks of its life. In the Serengeti, up to 90% of cheetah cubs are killed during this time by lions, leopards, hyenas, African Wild Dogs, or even by eagles. Cheetah cubs often hide in thick brush for safety. Mother cheetahs will defend their young and are at times successful in driving predators away from their cubs. Coalitions of male cheetahs can also chase away other predators, depending on the coalition size and the size and number of the predator. Because of its speed, a healthy adult cheetah has few enemies.
Relationship with humans
Cheetah fur was formerly regarded as a status symbol. Today, cheetahs have a growing economic importance for ecotourism and they are also found in zoos. White Oak Conservation in Yulee, Florida, which maintains a significant population of cheetahs, has cited that captive management presents challenges because of health, nutrition and socialization of the cats, but that these have been overcome through collaborations among wildlife facilities.
Cheetahs are far less aggressive than other felids, and in some parts of the world are considered a prestigious possession. Cheetah cubs are taken from the wild for the illegal wildlife trade and can be found for sale through street markets and the Internet. However, cheetahs don’t breed well in captivity and legal breeding facilities are unable to meet the demand. Thus, the proliferation of wild cheetah taken for the illegal pet trade has the potential of decimating wild populations. The Cheetah Conservation Fund estimates that hundreds of cheetah cubs are taken from the wild every year to be sold as pets; only about one in six survive.
Cheetahs living outside of protected areas often inhabit farmland, where they are shot by farmers who believe that they eat livestock. Recent evidence has shown, however, that cheetahs will not attack and eat livestock if they can avoid doing so, as they prefer their wild prey. The Cheetah Conservation Fund has designed and implemented programs to prevent predators' conflict with humans. These programs aim at helping the farmers to protect their livelihoods through education, livelihood development, habitat restoration and predator-friendly farming techniques, such as the highly-successful use of Livestock Guarding Dogs.
Ancient Egyptians often kept cheetahs as pets, and also tamed and trained them for hunting, although they did not domesticate it. Cheetahs would be taken to hunting fields in low-sided carts or by horseback, hooded and blindfolded, and kept on leashes while dogs flushed out their prey. When the prey was near enough, the cheetahs would be released and their blindfolds removed. This tradition was passed on to the ancient Persians and brought to India, where the practice was continued by Indian princes into the twentieth century. Cheetahs continued to be associated with royalty and elegance, their use as pets spreading just as their hunting skills were. Other such princes and kings kept them as pets, including Genghis Khan and Charlemagne, who boasted of having kept cheetahs within their palace grounds. Akbar the Great, ruler of the Mughal Empire from 1556 to 1605, kept as many as 1,000 cheetahs. As recently as the 1930s, the Emperor of Ethiopia, Haile Selassie, was often photographed leading a cheetah by a leash.
Cheetahs are still tamed in the modern world, much to their detriment as the demand in the illegal pet trade continues. However, their tameability has also allowed many registered institutions to educate the public by training cheetahs as educational ambassadors. One example is Burmani, who has been raised in England at Eagle Heights wild animal park from the age of three months. He was bred in a deer park in Germany. He is so tame that he has lost his hunting instinct.
Cheetah cubs have a high mortality rate due to predation by other carnivores, such as the lion and hyena, and perhaps genetic factors. It has been suggested that the low genetic diversity of cheetahs is a cause of poor sperm, birth defects, cramped teeth, kinked tails, and bent limbs. Some biologists even believe that they are too inbred to flourish as a species. Note, however, that they lost most of their genetic diversity thousands of years ago (see the beginning of this article), and yet have only been in decline in the last century or so, from 100,000 in the early 1900s to 10,000 today, due to loss of habitat and prey, human-wildlife conflict and the illegal pet trade.
Cheetahs are included on the International Union for Conservation of Nature (IUCN) list of vulnerable species (African subspecies threatened, Asiatic subspecies in critical situation) as well as on the US Endangered Species Act: threatened species - Appendix I of CITES (Convention on International Trade in Endangered Species). In 2014, the CITES Standing Committee recognized cheetahs as a ‘species of priority’ in north-east Africa in their strategies to counter wildlife trafficking.
Approximately 10,000 cheetahs remain in the wild in twenty-three African countries; Namibia has the most, with about 3,500. Another 50 to 60 critically endangered Asiatic cheetahs are thought to remain in Iran. There have been some successful breeding programs in zoos around the world. Additionally, recent research into improving in vitro fertilisation and embryo culture techniques have the potential of consistently producing embryos for transfer.
Founded in Namibia in 1990, the Cheetah Conservation Fund's mission is to be the world's resource charged with protecting the cheetah and to ensure its future. The organization works with all stakeholders within the cheetah's ecosystem to develop best practices in research, education and ecology and create a sustainable model from which all other species, including people, will benefit. The Cheetah Conservation Fund has close links and assists in training and sharing program successes with other countries where cheetahs live, including Botswana, Kenya, Tanzania, South Africa, Zimbabwe, Iran and Algeria. The organization's international program includes distributing materials, lending resources and support, and providing training through Africa and the rest of the world.
Re-introduction project in India
Asiatic cheetahs have been known to exist in India for a very long time, but as a result of hunting and other causes, cheetahs have been extinct in India since the 1940s. A captive propagation project has been proposed. Minister of Environment and Forests Jairam Ramesh told the Rajya Sabha on 7 July 2009, "The cheetah is the only animal that has been described extinct in India in the last 100 years. We have to get them from abroad to repopulate the species." He was responding to a call for attention from Rajiv Pratap Rudy of the Bharatiya Janata Party (BJP). "The plan to bring back the cheetah, which fell to indiscriminate hunting and complex factors like a fragile breeding pattern is audacious given the problems besetting tiger conservation." Two naturalists, Divya Bhanusinh and MK Ranjit Singh, suggested importing cheetahs from Africa, after which they will be bred in captivity and, in time, released in the wild.
However, the plan to reintroduce the African cheetahs to India has been suspended after discovering the distinctness between the cheetahs from Asia and Africa, having been separated between 32,000 to 67,000 years ago.
In popular culture
|This section does not cite any references or sources. (January 2013)|
- In Titian's Bacchus and Ariadne (1523), the god's chariot is borne by cheetahs (which were used as hunting animals in Renaissance Italy). Cheetahs were often associated with the god Dionysus, whom the Romans called Bacchus.
- George Stubbs' Cheetah with Two Indian Attendants and a Stag (1764–1765) also shows the cheetah as a hunting animal and commemorates the gift of a cheetah to George III by the English Governor of Madras, Sir George Pigot
- The Caress (1896), by the Belgian symbolist painter Fernand Khnopff (1858–1921), is a representation of the myth of Oedipus and the Sphinx and portrays a creature with a woman's head and a cheetah's body (often misidentified as a leopard's).
- André Mercier's Our Friend Yambo (1961) is a curious biography of a cheetah adopted by a French couple and brought to live in Paris. It is seen as a French answer to Born Free (1960), whose author, Joy Adamson, produced a cheetah biography of her own, The Spotted Sphinx (1969).
- Hussein, An Entertainment, a novel by Patrick O'Brian set in India of the British Raj period, illustrates the practice of royalty keeping and training cheetahs to hunt antelopes.
- The book How It Was with Dooms tells the true story of a family raising an orphaned cheetah cub named Duma (the Swahili word for cheetah) in Kenya. The films Cheetah (1989) and Duma (2005) were both loosely based on this book.
- Similarly, Roger Hunt successfully tames a cheetah in Willard Price's Safari Adventure, after rescuing it from an elephant pit trap. The cheetah soon befriends a German shepherd dog called Zulu.
- The animated series ThunderCats had a main character who was an anthropomorphic cheetah named Cheetara.
- In 1986, Frito-Lay introduced an anthropomorphic cheetah, Chester Cheetah, as the mascot for their Cheetos.
- Harold and Kumar Go to White Castle has a subplot involving an escaped cheetah, which later smokes cannabis with the pair and allows them to ride it.
- Comic book superheroine Wonder Woman's chief adversary is Dr. Barbara Ann Minerva, alias The Cheetah.
- On the CGI animated show Beast Wars: Transformers, Cheetor, one of the main characters on the Maximal faction, had the beast form of a cheetah. This was also carried over as the beast form of the Cheetor Hasbro transformer.
- The Japanese anime Damekko Dōbutsu features a clumsy but sweet-natured cheetah named Chiiko.
- The first release of Apple Inc.'s Mac OS X was code-named "Cheetah", which set the pattern for the subsequent releases being named after big cats.
- In Visionaries: Knights of the Magical Light the character Witterquick as the totem of a Cheetah and could turn into one.
- Titled "Hunting at 60 mph", the PlayStation 3 game Afrika features a Cheetah hunting a gazelle as the game's first "big game hunt".
- Wozencraft, W.C. (2005). "Order Carnivora". In Wilson, D.E.; Reeder, D.M. Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. pp. 532–533. ISBN 978-0-8018-8221-0. OCLC 62265494.
- Bauer, H., Belbachir, F., Durant, S., Hunter, L., Marker, L., Packer, K. & Purchase, N. (2008). "Acinonyx jubatus". IUCN Red List of Threatened Species. Version 2014.1. International Union for Conservation of Nature. Retrieved 29 August 2014.
- Sharp, N. C. (1997). "Timed running speed of a cheetah (Acinonyx jubatus)". Journal of Zoology, London 241 (3): 493–494. doi:10.1111/j.1469-7998.1997.tb04840.x.
- Milton Hildebrand (1959). "Motions of Cheetah and Horse". Journal of Mammalogy 40 (4): 481–495. doi:10.2307/1376265. JSTOR 1376265.[dead link] Although according to Cheetah, Luke Hunter and Dave Hamman, (Struik Publishers, 2003), pp. 37–38, the cheetah's fastest recorded speed was 110 km/h (68 mph).
- Carwardine, Mark (2008). Animal Records. New York: Sterling. p. 43. ISBN 978-1-4027-5623-8.
- Sears, Edward S. (2001). Running Through the Ages. Jefferson, North Carolina: McFarland. p. 5. ISBN 978-0-7864-0971-6.
- Smith, Roff (2 August 2012). "Cheetah Breaks Speed Record—Beats Usain Bolt by Seconds". National Geographic Daily News (National Geographic Society).
- "Speed sensation". Nature Video Collections. BBC Nature.
- "Cheetah fast facts". Zoological Society of London. Retrieved 5 June 2013.
- "Cheetah Fact Sheet" (PDF). Cheetah.org. Retrieved 3 June 2013.
- cheetah (n.d.). The American Heritage Dictionary of the English Language, Fourth Edition. Retrieved 2007-04-16.
- "Acinonyx jubatus". Cat Specialist Group. Retrieved 6 May 2014.
- O'Brien, S., D. Wildt, M. Bush (1986). "The Cheetah in Genetic Peril". Scientific American 254: 68–76.
- Young, T.P. and A.H. Harcourt (1997). "Viva Caughley!". Conservation Biology 11: 831–832. doi:10.1046/j.1523-1739.1997.011004831.x.
- Johnson, W. E., E. Eizirik, J. Pecon-Slattery, W. J. Murphy, A. Antunes, E. Teeling, S. J. O'Brien (2006). "The Late Miocene Radiation of Modern Felidae: A Genetic Assessment". Science 311 (5757): 73–77. doi:10.1126/science.1122277. PMID 16400146.
- Mattern, M. Y., D. A. McLennan (2000). "Phylogeny and Speciation of Felids". Cladistics 16 (2): 232–253. doi:10.1111/j.1096-0031.2000.tb00354.x.
- Wilson, D.E.; Reeder, D.M., eds. (2005). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. ISBN 978-0-8018-8221-0. OCLC 62265494.
- "Namibia: Cheetah Conservation Fund Celebrates 25 Years". allAfrica.com. 20 March 2015. Retrieved 6 April 2015.
- "Acinonyx jubatus (Cheetah, Hunting Leopard)". IUCNRedList.org. Retrieved 6 April 2015.
- Ella Davies (24 January 2011). "Iran's endangered cheetahs are a unique subspecies". Retrieved 6 April 2015.
- Three distinct cheetah populations, but Iran's on the brink, 18 January 2011, retrieved 6 April 2015
- "Cheetah, Acinonyx jubatus". Retrieved 6 April 2015.
- "Agility, Not Speed, Puts Cheetahs Ahead". Science 340: 1271. 14 June 2013. doi:10.1126/science.340.6138.1271-b.
- "Cheetah". Animal Info. 2006-03-21. Retrieved 2013-03-30.
- Boitani, Luigi, Simon & Schuster's Guide to Mammals. Simon & Schuster/Touchstone Books (1984), ISBN 978-0-671-42805-1
- "Cheetahs, Cheetah Pictures, Cheetah Facts - National Geographic". Animals.nationalgeographic.com. 2012-10-25. Retrieved 2013-03-30.
- Burnie D and Wilson DE (Eds.), Animal: The Definitive Visual Guide to the World's Wildlife. DK Adult (2005), ISBN 0-7894-7764-5
- "King Cheetah Fur Pattern Mutation". Cheetahspot.com. Retrieved 2013-03-30.
- "About the Cheetah". Retrieved 2015-06-03.
- "Dogs used to calm skittish cheetahs". DailyHerald.com. 2013-02-16. Retrieved 2013-03-30.
- Kaelin CB, Xu X, Hong LZ, David VA, McGowan KA, Schmidt-Küntzel A, Roelke ME, Pino J, Pontius J, Cooper GM, Manuel H, Swanson WF, Marker L, Harper CK, van Dyk A, Yue Bisong, Mullikin JC, Warren WC, Eizirik E, Kos L, O'Brien SJ, Barsh GS, Menotti-Raymond M (2012). "Specifying and Sustaining Pigmentation Patterns in Domestic and Wild Cats". Science 337 (6101): 1536–1541. doi:10.1126/science.1220893. PMID 22997338.
- "The lesser-spotted cheetah: Rare big cat without traditional markings sighted in wild for first time in nearly 100 years". Daily Mail (London). 2012-04-25. Retrieved 2013-03-06.
- "Asiatic Cheetah". Wild About Cats. Retrieved 2007-12-07.
- "Asiatic Cheetah". WWF-Pakistan. Retrieved 2007-12-07.
- "Scandal on the Serengeti: New light has been shed on the extent of female cheetahs' unfaithfulness to their male partners". inthenews.co.uk. May 30, 2007. Retrieved 2007-12-07.
- Eaton, Randall L. (1976) A Possible Case of Mimicry in Larger Mammals. Evolution 30(4):853-856 doi 10.2307/2407827
- , Cheetah cub survival revisited: a re-evaluation of the role of predation, especially by lions, and implications for conservation
- Richard Estes, foreword by Edward Osborne Wilson (1991) The Behavior Guide to African Mammals. University of California Press. Page 371.
- T. M. Caro (15 August 1994). Cheetahs of the Serengeti Plains: Group Living in an Asocial Species. University of Chicago Press. pp. 203–. ISBN 978-0-226-09433-5. Retrieved 21 March 2013.
- "Cheetah Information" (PDF). Cheetah Outreach. Retrieved 2015-03-06.
- MacMillan, Dianne. Cheetahs. Books.google.com. Retrieved 2013-03-30.
- Smithsonian Channel (October 24, 2014). "Cheetahs Fighting Cheetahs". youtube.com.
- Robert Eklund. "Eklund, Peters, Weise & Munro (2012)" (PDF). Roberteklund.info. Retrieved 2013-03-30.
- Robert Eklund. "Robert Eklund's Ingressive Phonation and Speech Page". Ingressivespeech.info. Retrieved 2013-03-30.
- Robert Eklund. "Robert Eklund's Homepage". Roberteklund.info. Retrieved 2013-03-30.
- Robert Eklund. "– The felid purring site". Purring.org. Retrieved 2013-03-30.
- Robert Eklund. "Eklund, Peters & Duthie (2010)" (PDF). Roberteklund.info. Retrieved 2013-03-30.
- Robert Eklund. "Eklund, Peters, Weise & Munro (2012)" (PDF). Roberteklund.info. Retrieved 2013-03-30.
- Robert Eklund. "Eklund & Peters (2013)" (PDF). roberteklund.info. Retrieved 2013-03-30.
- , Cheetah menu: wildlife instead of cattle .
- , Feeding ecology of the Asiatic cheetah Acinonyx jubatus venaticus in low prey habitats in northeastern Iran: Implications for effective conservation.
- Hunter, Luke & Hamman, Dave. Cheetah. Struik Publishers, 2003, p. 109.
- "Wild Cats of the World" - by Mel Sunquist and Fiona Sunquist
- Chinery, M. 1979. Killers of the wild: Streamlined cheetah. Wildlife 21:6-7.
- RVC Press Office (13 June 2013). "Groundbreaking RVC research shows wild cheetah reaching speeds of up to 58mph during a hunt". rvc.ac.uk. Royal Veterinary College, University of London. Retrieved 2013-06-16.
- Wilson AM, Lowe JC, Roskilly K, Hudson PE, Golabek KA, McNutt JW (2013). "Locomotion dynamics of hunting in wild cheetahs". Nature 498 (7453): 185–189. doi:10.1038/nature12295. PMID 23765495.
- "Cheetah tracking study reveals incredible acceleration". BBC News. 2013-06-12.
- Hunter, Luke & Hamman, Dave 2003, p. 96.
- Marker, L. Aspects of Namibian Cheetah (Acinonyx jubatus) Biology, Ecology and Conservation Strategies. PhD. Thesis, Department of Zoology, University of Oxford, United Kingdom. 2002.
- M. W. Hayward, M. Hofmeyr, J. O'Brien & G. I. H. Kerley (2006). "Prey preferences of the cheetah (Acinonyx jubatus) (Felidae: Carnivora): morphological limitations or the need to capture rapidly consumable prey before kleptoparasites arrive?". Journal of Zoology 270 (4): 615. doi:10.1111/j.1469-7998.2006.00184.x.
- "Cheetah". Retrieved 21 June 2013.
- Mitchell N., Tricorache P., Groom, R., Marker, L. and Durant, S. The Illegal Trade in Cheetahs. Poster. International Wildlife Trafficking Symposium. London, UK. February 2014.
- "Human Wildlife Conflict". Retrieved 2015-03-06.
- Fieldsports Britain. "Fieldsports Britain : Rabbits with a cheetah in Essex, grouse and". fieldsportschannel.tv. Retrieved 30 October 2012.
- Gugliotta, Guy (February 2008). "Rare Breed". Smithsonian Magazine. Retrieved 2008-03-07.
- "Cheetah Race for Survival". Cheetah.org. Retrieved 2015-03-06.
- Crosier, A.E., Henghali, J.N., Howard, J., Pukazhenthi, B.S., Terrell, K.A., Marker, L. and Wildt, D. Improved Quality of Cryopreserved Cheetah (Acinonyx jubatus) Spermatozoa After Centrifugation Through Accudenz. Journal of Andrology, Vol. 30, No. 3, May/June 2009
- The Times of India, Thursday, July 9, 2009, p. 11.
- "| Travel India Guide". Binoygupta.com. 18 May 2012. Retrieved 16 May 2013.
- "Breaking: India’s Plan to Re-Introduce the Cheetah on Hold". http://cheetah-watch.com. 8 May 2012. Retrieved 13 November 2014.
- Great Cats, Majestic Creatures of the Wild, ed. John Seidensticker, illus. Frank Knight, (Rodale Press, 1991), ISBN 0-87857-965-6
- Cheetah, Katherine (or Kathrine) & Karl Ammann, Arco Pub, (1985), ISBN 0-668-06259-2.
- Cheetah (Big Cat Diary), Jonathan Scott, Angela Scott, (HarperCollins, 2005), ISBN 0-00-714920-4
- Science (vol 311, p 73)
- Cheetah, Luke Hunter and Dave Hamman, (Struik Publishers, 2003), ISBN 1-86872-719-X
- Allsen, Thomas T. (2006). "Natural History and Cultural History: The Circulation of Hunting Leopards in Eurasia, Seventh-Seventeenth Centuries." In: Contact and Exchange in the Ancient World. Ed. Victor H. Mair. University of Hawai'i Press. Pp. 116–135. ISBN ISBN 978-0-8248-2884-4; ISBN ISBN 0-8248-2884-4
- Eklund, Robert & Gustav Peters. 2013. A comparative acoustic analysis of purring in juvenile, subadult and adult cheetahs. In: Robert Eklund (editor.), Proceedings of Fonetik 2013, the XXVIth Swedish Phonetics Conference, Studies in Language and Culture, no. 21, ISBN 978-91-7519-582-7, eISBN 978-91-7519-579-7, ISSN 1403-2570, pp. 25–28.
- Eklund, Robert, Gustav Peters & Elizabeth D. Duthie. 2010 (third edition). An acoustic analysis of purring in the cheetah (Acinonyx jubatus) and in the domestic cat (Felis catus). Proceedings of Fonetik 2010, Lund University, 2–4 June 2010, Lund, Sweden, pp. 17–22. Download from  or .
- Gus Mills, M. G. L. Mills, Martin Harvey (2005). African Predators. Struik. ISBN 1-77007-220-9.
- Gus Mills (1998). Big Cats and Other African Carnivores. Struik. ISBN 1-86825-920-X.
- Caro, T. M. (1994). Cheetahs of the Serengeti Plains: group living in an asocial species. Chicago: University of Chicago Press. ISBN 0-226-09433-2.
|Wikimedia Commons has media related to Acinonyx jubatus.|
|Wikispecies has information related to: Acinonyx jubatus|
- Species portrait Cheetah; IUCN/SSC Cat Specialist Group
- Cheetah at the Encyclopedia of Life
- Biodiversity Heritage Library bibliography for Acinonyx jubatus
- Cheetah Conservation Fund
- Save China's Tigers to Fund Wild Cat Conservation Worldwide
- De Wildt Cheetah and Wildlife Trust
- On the Chase With Cheetahs - slideshow by Life magazine
- Fake Flies and Cheating Cheetahs: measuring the speed of a cheetah
- Mutant Cheetahs: information on color variants of cheetahs
- 110km/h Cheetah attack gazelle Video showing cheetah's speed, running mechanics, and hyenas stealing a cheetah's prey.